The leaf whorls are placed in the genus _Annularia_. One form commonly
found in Illinois has long, pointed, needlelike leaves and is given the
name _Asterophyllites_. _Calamostachys_, shown on plate 5, is one of the
most common calamite cones.
SPHENOPHYLLUM
(Plate 2)
The name _Sphenophyllum_ refers to both stems and leaves of this extinct
genus, which was related to the scouring rushes—note its resemblance to
_Annularia_.
A small herbaceous plant, _Sphenophyllum_ formed much of the swampy
undergrowth of the Pennsylvanian Period and is abundant among Illinois
fossils. It had a slender, ribbed stem bearing whorls of delicate,
wedge-shaped leaves, generally less than three-fourths of an inch long,
attached around the stem in multiples of three.
The cones of this group also are slender, delicate structures, bearing a
number of sporangia, and are correctly called _Bowmanites_, although
they also have been called _Sphenophyllostachys_. These fossil cones
frequently are found in Illinois.
_Sphenophyllum_ first appeared during the Devonian Period, some 300
million years ago, but did not become abundant until Pennsylvanian time.
The genus continued through the Permian but died out in Triassic time.
FERNS
(Plates 1 and 3)
True ferns, like those living in today’s woodlands, were common in the
Pennsylvanian forests. Some species attained heights of 30 to 40 feet.
Their fronds (compound leaves divided into segments or leaflets)
commonly were five to six feet long.
[Illustration: Portion of fern frond showing sori on lower side of
leaflets]
True ferns do not produce cones or seeds, but spores, which develop in
cases called sporangia. The sporangia frond showing are attached in
clusters (sori) to the lower side or margins of the leaves. In modern
ferns the sporangia may also occur on fertile spikes.
The shape and position of the sori are used to identify modern ferns,
but because leaves that bear sori (“fertile” leaves) are rare among
fossil specimens, the number, shape, and attachment of the leaflets and
the pattern of the veins are more commonly used for identification.
[Illustration: Reconstruction of _Megaphyton_
(after Hirmer)]
Public-domain text, read in full here on John Shaqi.
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