=Wood Strand in Monocotyledons and Dicotyledons.=--Each wood strand (or
fibro-vascular bundle) consists of two parts--the bast and the wood
proper. The wood is on the side of the strand toward the center of the
stem and contains large tubular canals that take the watery sap upward
from the roots. The bast is on the side toward the bark and contains
fine tubes through which diffuses the dense sap containing digested
food from the leaves. In the root (Fig. 71) the bast and the wood are
separate, so that there are _two kinds of strands_.
[Illustration: FIG. 72.--PART OF CROSS-SECTION OF ROOTSTOCK OF
ASPARAGUS, showing a few fibro-vascular bundles. An endogenous stem.]
In monocotyledons, as already said, the strands (or bundles) _are
usually scattered in the stem with no definite arrangement_ (Figs. 72,
73). In dicotyledons the strands, or bundles, _are arranged in a ring_.
As the dicotyledonous seed germinates, five bundles are usually formed
in its hypocotyl (Fig. 74); soon five more are interposed between them,
and the multiplication continues, in tough plants, until the bundles
touch (Fig. 74, right). The inner parts thus form a ring of wood and
the outer parts form the inner bark or bast. A new ring of wood or bast
is formed on stems of dicotyledons each year and the age of a cut stem
is easily determined.
[Illustration: _a_
_b_
FIG. 73.--THE SCATTERED BUNDLES OR STRANDS, in monocotyledons at _a_,
and the bundles in a circle in dicotyledons at _b_.]
[Illustration: FIG. 74.--DICOTYLEDONOUS STEM OF ONE YEAR AT LEFT WITH
FIVE BUNDLES, and a Two-year Stem at Right.
_o_, the pith; _c_, the wood part; _b_, the bast part; _a_, one year’s
growth.]
[Illustration: FIG. 75.--FIBRO-VASCULAR BUNDLE OF INDIAN CORN, much
magnified.
_A_, annular vessel; _A′_, annular or spiral vessel; _TT′_,
thick-walled vessels; _W_, tracheids or woody tissue; _F_, sheath of
fibrous tissue surrounding the bundle; _FT_, fundamental tissue or
pith; _S_, sieve tissue; _P_, sieve plate; _C_, companion cell; _l_,
intercellular space, formed by tearing down of adjacent cells; _W′_,
wood parenchyma.]
[Illustration: FIG. 76.--THE DICOTYLEDONOUS BUNDLE OR WOOD STRAND.
Upper figure is of moonseed:
_c_, cambium; _d_, ducts; 1, end of first year’s growth; 2, end of
second year’s growth; bast part at left and wood part at right. Lower
figure (from Wettstein) is sunflower; _h_, wood-cells; _g_, vessels;
_c_, cambium; _p_, fundamental tissue or parenchyma; _b_, bast; _bp_,
bast parenchyma; _s_, sieve-tubes.]
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