When cross-sections of monocotyledonous and dicotyledonous bundles are
examined under the microscope, it is readily seen why dicotyledonous
bundles form rings of wood and monocotyledonous cannot (Figs. 75
and 76). The dicotyledonous bundle (Fig. 76) has, running across
it, a layer of brick-shaped cells called =cambium=, which cells are
a specialized form of the parenchyma cells and retain the power of
growing and multiplying. The bundles containing cambium are called
_open bundles_. There is no cambium in monocotyledonous bundles (Fig.
75) and the bundles are called _closed bundles_. Monocotyledonous stems
_soon cease to grow in diameter_. The stem of a palm tree is almost as
large at the top as at the base. As dicotyledonous plants grow, the
_stems become thicker each year_, for the delicate active cambium layer
forms new cells from early spring until midsummer or autumn, adding to
the wood within and to the bark without. As the growth in spring is
very rapid, the first wood-cells formed are much larger than the last
wood-cells formed by the slow growth of the late season, and the spring
wood is less dense and lighter colored than the summer wood; hence the
time between two years’ growth is readily made out (Figs. 77 and 78).
Because of the rapid growth of the cambium in spring and its consequent
soft walls and fluid contents, the bark of trees “peels” readily at
that season.
[Illustration: FIG. 77.--WHITE PINE STEM, 5 years old. The outermost
layer is bark.]
=Medullary Rays.=--The first year’s growth in dicotyledons forms a
woody ring which almost incloses the pith, and this is left as a small
cylinder which does not grow larger, even if the tree should live a
century. It is not quite inclosed, however, for the narrow layers
of soft cells separating the bundles remain between them (Fig. 78),
forming radiating lines called =medullary rays= or =pith rays=.
[Illustration: FIG. 78.--ARRANGEMENT OF TISSUES IN TWO-YEAR-OLD STEM OF
MOONSEED.
_p_, pith; _f_, parenchyma. The fibro-vascular bundles, or wood
strands, are very prominent, with thin medullary rays between.]
=The Several Plant Cells and their Functions.=--In the =wood= there are
some parenchyma cells that are still with thin walls, but have lost
the power of division. They are now _storage cells_. There are also
wood fibers which are thick-walled and rigid (_h_, Fig. 76), and serve
to _support_ the sap-canals or _wood vessels_ (or tracheids) that are
formed by the absorption of the end walls of upright rows of cells; the
canals pass from the roots to the twigs and even to ribs of the leaves
and serve to transport the root water. They are recognized (Fig. 79) by
the peculiar thickening of the wall on the inner surface of the tubes,
occurring in the form of spirals. Sometimes the whole wall is thickened
except in spots called _pits_ (_g_, Fig. 76). These thin spots (Fig.
80) allow the sap to pass to other cells or to neighboring vessels.
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