[Illustration: FIG. 79.--MARKINGS IN CELL WALLS OF WOOD FIBERS.
_sp_, spiral; _an_, annular; _sc_, scalariform.]
[Illustration: FIG. 80.--PITS IN THE CELL WALL.
Longitudinal section of wall at _b_, showing pit borders at _o_, _o_.]
=The cambium=, as we have seen, consists of cells whose function is
_growth_. These cells are thin-walled and filled with protoplasm.
During the growing season they are continually adding to the wood
within and the bark without; hence the layer moves outward as it
deposits the new woody layer within.
[Illustration: FIG. 81.--SIEVE-TUBES, _s_, _s_;
_p_ shows a top view of a sieve-plate, with a companion cell. _c_, at
the side; _o_ shows sieve-plates in the side of the cell. In _s_, _s_
the protoplasm is shrunken from the walls by reagents.]
[Illustration: FIG. 82.--THICK-WALLED BAST CELLS.]
=The bark= consists of inner or _fibrous bark_ or new bast (these
fibers in flax become linen), the _green or middle bark_ which
functions somewhat as the leaves, and the _corky or outer bark_. The
common word “bark” is seen therefore not to represent a homogeneous or
simple structure, but rather a collection of several kinds of tissue,
all separating from the wood beneath by means of cambium. The new
bast contains (1) the _sieve-tubes_ (Fig. 81) which transport the sap
containing organic substances, as sugar and proteids, from the leaves
to the parts needing it (_s_, Fig. 76). These tubes have been formed
like the wood vessels, but they have sieve-plates to allow the dense
organic-laden sap to pass with sufficient readiness for purposes of
rapid distribution. (2) There are also thick-walled _bast fibers_
(Fig. 82) in the bast that serve for _support_. (3) There is also
some parenchyma (parent tissue) in the new bast; it is now in part a
_storage_ tissue. Sometimes the walls of parenchyma cells in the cortex
thicken at the corners and form _brace cells_ (Fig. 83) (collenchyma)
for _support_; sometimes the whole wall is thickened, forming _grit
cells_ or _stone cells_ (Fig. 84; examples in tough parts of pear, or
in stone of fruits). Some parts serve for secretions (milk, rosin,
etc.) and are called _latex tubes_.
[Illustration: FIG. 83.--COLLENCHYMA IN WILD JEWELWEED OR TOUCH-ME-NOT
(IMPATIENS).]
[Illustration: FIG. 84.--GRIT CELLS.]
=The outer bark= of old shoots consists of _corky_ cells that _protect_
from mechanical injury, and that contain a fatty substance (suberin)
impermeable to water and of service to _keep in moisture_. There is
sometimes a cork cambium (or phellogen) in the bark that serves to
extend the bark and keep it from splitting, thus increasing its power
to protect.
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