Form and Function: A Contribution to the History of Animal Morphology — John Shaqi
Form and Function: A Contribution to the History of Animal MorphologyRussell, E. S. (Edward Stuart)
History
Form and Function: A Contribution to the History of Animal Morphology
Russell, E. S. (Edward Stuart)
Anatomy, Comparative -- History; Morphology (Animals) -- History
We may conveniently refer here to one or two other ideas of Aristotle
regarding the causes of form. He makes the profound remark that the
possible range of form of an organ is limited to some extent by its
existing differentiation. Thus he explains the absence of external
(projecting) ears in birds and reptiles by the fact that their skin is
hard and does not easily take on the form of an external ear (_De
Partibus_, ii, 12). The fact of the inverse correlation is certain;
the explanation is, though very vague, probably correct.
In one passage of the _De Partibus_ Aristotle clearly enunciates the
principle of the division of labour, afterwards emphasised by H.
Milne-Edwards. In some insects, he says, the proboscis combines the
functions of a tongue and a sting, in others the tongue and the sting
are quite separate. "Now it is better," he goes on, "that one and the
same instrument shall not be made to serve several dissimilar ends;
but that there shall be one organ to serve as a weapon, which can then
be very sharp, and a distinct one to serve as a tongue, which can then
be of spongy texture and fit to absorb nutriment. Whenever, therefore,
Nature is able to provide two separate instruments for two separate
uses, without the one hampering the other, she does so, instead of
acting like a coppersmith who for cheapness makes a spit and
lampholder in one" (iv., 6, 683^a).
(5) The first sentence of the _Historia Animalium_ formulates, with
that simplicity and directness which is so characteristic of
Aristotle, the distinction between homogeneous and heterogeneous
parts, in the mass the distinction between tissues and organs. "Some
parts of animals are simple, and these can be divided into like parts,
as flesh into pieces of flesh; others are compound, and cannot be
divided into like parts, as the hand cannot be divided into hands, nor
the face into faces. All the compound parts also are made up of simple
parts--the hand, for example, of flesh and sinew and bone" (Cresswell,
_loc. cit_., p. I).
In the _De Partibus Animalium_ he broadens the conception by adding
another form of composition. "Now there are," he says, "three degrees
of composition; and of these the first in order, as all will allow, is
composition out of what some call the elements, such as earth, air,
water, fire.... The second degree of composition is that by which the
homogeneous parts of animals, such as bone, flesh, and the like, are
constituted out of the primary substances. The third and last stage is
the composition which forms the heterogeneous parts, such as face,
hand, and the rest" (ii., 1, 646^a, trans. Ogle).
Public-domain text, read in full here on John Shaqi.
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