Fossil plants, Vol. 2 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 2 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
The recent members of the Lycopodiales are considered apart from
the extinct genera in order that our examination of the latter may
be facilitated by a knowledge of the salient characteristics of the
surviving types of this important section of the Pteridophyta. A
general acquaintance with the extinct as well as with the recent genera
will enable us to appreciate the contrasts between the living and the
fossil forms and to realise the prominent position occupied by this
group in the Palaeozoic period, a position in striking contrast to
the part played by the diminutive survivors in the vegetation of the
present day. In the account of the recent genera special attention is
drawn to such features as afford a clue to the interpretation of the
fossils, and the point of view adopted, which at times may appear to
lead to an excessive attention to details, is necessarily somewhat
different from that represented in botanical text-books[83].
A. =HOMOSPOREAE.=
=Lycopodiaceae=: genera _Phylloglossum_, _Lycopodium_.
B. =HETEROSPOREAE.=
=Selaginellaceae=: genus _Selaginella_.
=Isoetaceae=: genus _Isoetes_.
The existing plants included in the Lycopodiales are in nearly all
cases perennial herbaceous pteridophytes, exhibiting in their
life-histories a well marked alternation of generations. The sporophyte
(asexual generation) is characterised by the relatively small size
of the leaves except in the genus _Isoetes_ (fig. 132) and in the
Australian and New Zealand genus _Phylloglossum_. The stems are usually
erect or trailing, pendulous in epiphytic species or small and tuberous
in _Isoetes_ and _Phylloglossum_. The repeated forking of the shoots
(monopodial and dichotomous branching) is a prominent feature of the
group. The vascular tissue of the stem usually assumes the form of a
single axial strand (stele) (fig. 125), but the shoots of some species
of _Selaginella_ often contain two or more distinct steles (fig. 131).
The group as a whole is characterised by the centripetal development of
the xylem composed almost entirely of scalariform tracheids: secondary
xylem and phloem of a peculiar type occur in _Isoetes_, and the
production of secondary xylem elements in a very slight degree has been
noticed in one species of _Selaginella_ (_S. spinosa_)[84]. The roots
are constructed on a simple plan, having in most cases only one strand
of spiral protoxylem elements (monarch structure). In _Lycopodium_,
in which stem and root anatomy are more nearly of the same type than
in the majority of plants, several protoxylem strands may be present.
The sporangia are axillary or, more frequently, borne on the upper
surface of sporophylls, which are either identical with or more or less
distinct from the foliage leaves; in the latter case the sporophylls
often occur in the form of a well defined strobilus (cone) at the tips
of branches.
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