Fossil plants, Vol. 2 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 2 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
The gametophyte (sexual generation) is represented by prothalli which,
in the homosporous genera, may live underground as saprophytes, or the
upper portion may develop chlorophyll and project above the surface of
the ground as an irregularly lobed green structure (_e.g._ _Lycopodium
cernuum_)[85]. In the heterosporous forms the prothalli are much
reduced and do not lead an independent existence outside the spore by
the membrane of which they are always more or less enclosed. The sexual
organs are represented by antheridia and archegonia; the male cells
are provided with two cilia except in _Isoetes_ which has multiciliate
antherozoids like those of the ferns.
The existing Lycopods, though widely distributed, never grow in
sufficiently dense masses to the exclusion of other plants to form a
conspicuous feature in the vegetation of a country. The inconspicuous
rôle which they play among the plant-associations of the present era
affords a striking contrast to the abundance of the arborescent species
in the Palaeozoic forests of the northern hemisphere.
• • • • •
=Lycopodiaceae.= _Lycopodium_, represented by nearly 100 species,
forms a constituent of most floras: epiphytic species predominate in
tropical regions, while others flourish on the mountains and moorlands
of Britain and in other extra-tropical countries. For the most part
_Lycopodium_ exhibits a preference for a moist climate and appears to
be well adapted to habitats where the amount of sunlight is relatively
small and the conditions of life unfavourable for dense vegetation.
Mountains and islands constantly recur as situations from which species
have been recorded. Some species are essentially swamp-plants, _e.g._
_Lycopodium inundatum_, a British species, and _L. cruentum_ from
the marshes of Sierra Nevada. A variety of the American species, _L.
alopecuroides_ (var. _aquaticum_) affords an instance of a submerged
form, which has been collected from an altitude of 12–14,000 ft. on the
Andes and Himalayas. It is noteworthy that a considerable variety of
habitats is represented within the limits of the genus and that many
species are sufficiently hardy to exist in circumstances which would be
intolerable to the majority of flowering plants[86].
The British species frequently spoken of as Club Mosses, include
_Lycopodium Selago_, _L. annotinum_, _L. clavatum_, _L. alpinum_, and
_L. inundatum_.
• • • • •
=Selaginellaceae.= The species of _Selaginella_, over 300 in number,
are widely spread in tropical and subtropical forests, growing on the
ground with trailing, suberect or erect stems climbing over taller and
stouter plants or as pendulous epiphytes on forest trees.
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