Certain problematical fossils found in association with the sterile leaves of _Dolerophyllum Berthieri_ have been described by Renault as the male organs of that species. These are elliptical discs, 6 × 5 cm., with an excentrically placed stalk: embedded in a carbonised lamina are numerous rows of elliptical bodies, 410μ × 280μ, characterised by two curved longitudinal grooves on the surface and regarded by Renault as pollen-grains. The chains of these microspores radiate outwards from the neighbourhood of the stalk and cover most of the surface of the disc (fig. 432, B). Some silicified pieces of similar spore-bearing discs from Grand’ Croix named _Dolerophyllum fertile_[390] afford additional information as to these remarkable reproductive organs. The earlier account of this species by Renault is confirmed by Solms-Laubach[391] who examined the original sections. The peltate fleshy discs preserved as incomplete specimens consist of lacunar parenchyma 15–18 mm. thick traversed at right angles to the surface by numerous loculi (fig. 432, C), circular or oval in transverse section, containing large numbers of microspores, _s_, similar in size and form to those on the carbonised discs of the Mt Pelé specimen. Vascular strands occur between and parallel to the spore-chambers. The spores contain 8–10 cells (fig. 432, D) and Renault believes that dehiscence of the exine occurred along the two deep grooves which mark the limits of an operculum. He emphasises the peculiar structure of the microspores by speaking of them as prepollinia: in size and in the presence of internal cells (? male prothallus) they resemble the spores found in the pollen-chamber of a seed described by Renault as _Aetheotesta elliptica_[392] which he thinks may belong to a member of the Dolerophylleae. It has also been suggested that _Codonospermum_ may be a seed of _Dolerophyllum_[393]. An unconvincing specimen described by Saporta and Marion[394] as a seed-bearing bract is regarded by them as referable to _Dolerophyllum_, but the evidence for any connexion is far from satisfactory.