There is nothing definite to be said with regard to the affinity of _Dolerophyllum Goepperti_ or the microsporophylls represented by _D. fertile_ and the specimens associated with _D. Berthieri_. Renault considers that both sterile and fertile specimens belong to the same genus, which he assigns to a position between Pteridophytes and Cycads. As Solms-Laubach says, the evidence supplied by the structure of the veins of _D. Goepperti_ in favour of a cycadean alliance is not convincing. The type of vernation is unlike that of any known Cycad or indeed of any Gymnosperm: the large size of the leaves is another though weaker objection to this comparison, as the pinnae of _Bowenia_ (fig. 391) and especially those of some species of _Zamia_ (fig. 388), are of equal or larger dimensions. If, as seems probable, the xylem-strands are mesarch that is a point of contact with recent Cycads, but the bundle as a whole bears but a remote resemblance to that of a cycadean leaf and is much more like the veins of _Ginkgo_. The bud shown in fig. 430, A, is probably a young shoot and not merely a large compound leaf. If it were an unexpanded frond of _Neuropteris_ bearing _Cyclopteris_ pinnules we should expect to find indications of scattered desmogen-strands such as would occur in the _Myeloxylon_ type of rachis. The resemblance to most forms of _Cordaites_ is by no means close though a few leaves referred to that genus (_e.g._ _C. circularis_, fig. 468, B) are similar to those of _Dolerophyllum_[395]. The male organs are unlike those of any other plant: they may be described as sporophylls with microsporangia or perhaps synangia embedded in the mesophyll and containing microspores similar to those of some Pteridosperms or true Gymnosperms. Attention has been called to the close resemblance of the spores shown in fig. 432, C, D to those recently discovered by Kidston and referred to the genus _Whittleseya_ (fig. 429), and it is very probable that the striking similarity is an index of affinity. =Ottokaria.= Zeiller.