_Zooecia._ Although the zooecia are distinctly tubular as a whole, two
longitudinal axes may be distinguished in each, for the tip is bent
upwards in a slanting direction, bearing the orifice at its extremity.
The main axis is, however, at right angles to the dorso-ventral axis,
and the dorsal surface, owing to the position of the aperture, can
always be readily distinguished from the ventral, even when the position
of the zooecium is vertical. Each zooecium tapers towards the posterior
extremity. Parietal muscles are always present.
[Illustration: Fig. 36.--Structure of _Paludicella ehrenbergi_ (A and B
after Allman).
A=a single zooecium with the polypide retracted. B=the base of the
lophophore as seen from above with the tentacles removed. C=the orifice
of a polypide with the collar expanded and the tentacles partly
retracted. _a_=tentacles; _c_=collar; _d_=mouth; _e_=oesophagus;
_f_=stomach; _g_=intestine; _k_=parieto-vaginal muscles; _p_=parietal
muscles; _o_=cardiac part of the stomach; _r_=retractor muscle;
_s_=funiculus.]
_Polypide._ The most striking features of the polypide are the absence
of any trace of a gizzard and the highly specialized form assumed by the
cardiac part of the stomach. There are two funiculi, both connecting the
pyloric part of the stomach with the endocyst. The ovary develops at the
end of the upper, the testis at that of the lower funiculus.
_Resting buds._ The resting buds are spindle-shaped.
Kraepelin recognized two species in the genus mainly by their method of
growth and the number of tentacles. In his _P. mülleri_ the zoarium is
always recumbent and the polypide has 8 tentacles, whereas in _P.
articulata_ or _ehrenbergi_ the tentacles number 16 and upright branches
are usually developed. It is probable, however, that the former species
should be assigned to _Victorella_, for it is often difficult to
distinguish _Paludicella_ from young specimens of _Victorella_ unless
the latter bear adventitious terminal buds. The gizzard of _Victorella_
can be detected in well-preserved material even under a fairly low power
of the microscope, and I have examined specimens of what I believe to be
the adult of _mülleri_ which certainly belong to that genus.
Public-domain text, read in full here on John Shaqi.
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