Fungi: Their Nature and UsesCooke, M. C. (Mordecai Cubitt)
Science
Fungi: Their Nature and Uses
Cooke, M. C. (Mordecai Cubitt)
Fungi
Much interest attaches to the other sub-family of _Coniomycetes_, in which
the species are produced for the most part on living plants. So much
has been discovered during recent years of the polymorphism which
subsists amongst the species in this section, that any detailed
classification can only be regarded as provisional. Hence we shall
proceed here upon the supposition that we are dealing with autonomous
species. In the first place, we must recognize a small section in which
a kind of cellular peridium is present. This is the _Æcidiacei_, or order
of "cluster cups." The majority of species are very beautiful objects
under the microscope; the peridia are distinctly cellular, and white or
pallid, produced beneath the cuticle, through which they burst, and,
rupturing at the apex, in one genus in a stellate manner, so that the
teeth, becoming reflexed, resemble delicate fringed cups, with the
orange, golden, brown, or whitish spores or pseudospores nestling in the
interior.[J] These pseudospores are at first produced in chains, but
ultimately separate. In many cases these cups are either accompanied or
preceded by spermogonia. In two other orders there is no peridium. In the
_Cæomacei_, the pseudospores are more or less globose or ovate, sometimes
laterally compressed and simple; and in _Pucciniæi_, they are elongated,
often subfusiform and septate. In both, the pseudospores are produced
in tufts or clusters _direct from the mycelium. The Cæomacei_ might
again be subdivided into _Ustilagines_[K] and _Uredines_.[L] In the
former, the pseudospores are mostly dingy brown or blackish, and in the
latter more brightly coloured, often yellowish. The _Ustilagines_
include the smuts and bunt of corn-plants, the _Uredines_ include the
red rusts of wheat and grasses. In some of the species included in the
latter, two forms of fruit are found. In _Melampsora_, the summer
pseudospores are yellow, globose, and were formerly classed as a species
of _Lecythea_, whilst the winter pseudospores are brownish, elongated,
wedge-shaped by compression, and compact. The _Pucciniæi_[M] differ
primarily in the septate pseudospores, which in one genus (_Puccinia_) are
uniseptate; in _Triphragmium_, they are biseptate; in _Phragmidium_,
multiseptate; and in _Xenodochus_, moniliform, breaking up into
distinct articulations. It is probable that, in all of these, as is
known to be the case in most, the septate pseudospores are preceded or
accompanied by simple pseudospores, to which they are mysteriously
related. There is still another, somewhat singular, group usually
associated with the _Pucciniæi_, in which the septate pseudospores are
immersed in gelatin, so that in many features the species seem to
approach the _Tremellini_. This group includes two or three genera, the
type of which will be found in _Podisoma_.[N] These fungi are parasitic on
living junipers in Britain and North America, appearing year after year
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