It is of interest to consider the general characteristics of the members
of the Fern tribe. As a rule the stem is either in the nature of a short
underground process bearing a rosette of leaves, as in the case of the
Male Fern and Hartstongue, or there is a horizontal stem more or less
below the surface of the soil, such as is to be seen in the case of the
Bracken Fern and the Polypody. Sometimes the stem assumes the proportions
of a trunk, but these Tree Ferns only occur in the tropics. Where the
stem of the Fern is upright it is properly termed a caudex, whilst in its
horizontal form it is spoken of as a rhizome. There is actually some
doubt as to the real nature of the frond of the Fern. Some botanists are
inclined to believe that it is not really a leaf at all, but is a
modified stem structure. Those who hold this view consider that the
curious scaly structures so common amongst Ferns are really the leaves of
the plant. Here the matter must be left on the present occasion, as it is
proposed to use the terms leaf and frond as meaning the same thing.
An outstanding feature in the case of most Ferns is the remarkable manner
in which the fronds are subdivided. In the case of the Male Fern it is
seen that the upper part of the stalk, or rachis, as it is called, bears
two rows of leaflets. These leaflets are properly referred to as pinnæ.
When the leaflets are subdivided the divisions are spoken of as pinnules.
These pinnules may be deeply lobed, and when this is the case each lobe
is called a segment. In very large fronds the pinnules are again divided;
the frond is then said to be tri-pinnate. Sometimes towards the top of
the pinnæ or the frond the divisions become less pronounced; this
character is designated pinnatifid. It should be noticed that the lower
portion of the stalk, on which there are no pinnæ, is called a stipes. Of
course in some cases, as with the Hartstongue, the leaf is quite
undivided, without even any very pronounced indentations on the margin.
The unrolling of the Fern frond is a very beautiful process. Where the
leaf is not divided in any way the process of expanding resembles the
uncoiling of a watch-spring. Even where there are divisions the unrolling
goes forward in the same manner with each subdivision, even down to the
lobes. This particular mode of unfolding is called circinate. The texture
of the leaves of Ferns is mostly thin and delicate, so that apart from
some exceptions the foliage is not able to withstand the action of dry
air. A notable feature with a large number of Ferns is the length of time
which the leaves take to develop. The fronds of the Male Fern, for
instance, start in the bud at least two years before they actually
unfold. An examination will show that the roots of the Male Fern spring
from the frond bases. It will be found that the position of the roots is
the same in all Ferns.
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