With all Ferns the production of spores is confined to the leaves. In
many instances there is no distinction between the fertile and the barren
leaves. The stem does not start at once to produce leaves bearing the
sporangia or spore cases. Thus, in the very young Fern the fronds are
always barren; as the stem becomes older, fertile fronds are produced. In
some cases the sporangia are borne on distinct leaves, as in the case of
the Hard Fern, or on special parts of the leaves, in the manner to be
seen in the Royal Fern. The difference in such cases is not really a very
important distinction. A careful examination of the fertile portion of a
Royal Fern frond will show a small amount of green tissue, or mesophyll,
as it is called, at the lower portion of the pinnæ. Actually the fertile
leaf, or part of a leaf, is similar to the barren portions, save that it
produces a much reduced amount of green tissue or, in some cases, perhaps
none at all.
In general appearance the Club Mosses bear a resemblance to the true
Mosses, and hence the popular name, which is certainly rather misleading.
With these plants the leaves are small and almost bristle-like, and are
gathered closely round the stem. In many of the Club Mosses a large part
of the stem lies closely along the ground, and from this at intervals
roots are sent down into the soil and leafy shoots rise upwards. The
sporangia are produced on special leaves, which are usually gathered
together in the form of cones.
Although they vary somewhat in size, all the Horsetails are striking
plants. Here there is a branching underground rhizome from which arise
the aerial stems. The most distinctive feature of the plant are the
whorls of smaller branches which arise from the joints of the main stem.
These carry on the work which is usually assigned to the foliage of the
average plant,—that is, the assimilating of carbon from the carbonic acid
of the atmosphere. The real leaves of the Horsetail are much reduced in
size, and take very little part in the work of nutrition. We shall find
them at the joints of the stem as rings, each collection forming a kind
of sheath. The leaves, which are usually of the same number as the
branches, show no sign of their individuality, save in the little
projecting teeth. In some species the fertile shoots, which appear in the
form of cones, are produced specially. These appear in the spring before
the ordinary vegetative growths, and are quite destitute of chlorophyll.
In other species the normal green shoots are fertile at the termination.
The sporangia are borne on curious scale-like leaves, a large number of
which go to the making of a cone.
Public-domain text, read in full here on John Shaqi.
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