The same series of phenomena may be shown in another way. If
a contractile strip of tolerable length be obtained, with the
waves passing freely from one end to the other, and if a series
of parallel and equidistant cuts be made along one side of the
strip, in a direction at right angles to the length, and each cut
extending two-thirds of the breadth of the strip, the chances are
in favour of the contraction-waves being wholly unaffected by the
sections, however numerous these may be. But now, if another series
of parallel and equidistant cuts of the same length as the first
ones, and alternating with them, be made along the other side
of the contractile strip, the result is, of course, a number of
interdigitating cuts; and it is easy to see that by beginning with a
few such cuts and progressively increasing their number, a point must
somewhere be reached at which one portion will become physiologically
separated from the rest. The amount of such section, however, which
contractile strips will sometimes endure is truly surprising. I
have seen such a strip twenty inches long by one and a half inches
wide with ten such cuts along each side, and the contraction-waves
passing without impediment from end to end. But what I wish more
especially to observe just now is, that by progressively increasing
the number of such interdigitating cuts up to the point at which the
contraction-wave is blocked, and then leaving the tissue to recover
itself, in many cases it will be observed that the blocking is sooner
or later overcome; that on then adding more interdigitating cuts the
blocking again supervenes; but that in time it may again be overcome,
and so on. It is, however, comparatively rare to find cases in which
blocking is overcome twice or thrice in succession.
Section is not the only way in which blocking of waves may be caused
in contractile strips. I find that pressure, even though very gentle,
exerted on any part of a strip causes a blocking of the waves at that
part, even after the pressure has been removed. If the pressure has
been long continued, after its removal the blocking will probably
be permanent; but if the pressure has been only of short duration,
the blocking will most likely be transitory. Even the slight strains
caused by handling a contractile strip in the air are generally
followed by a decrease in the rate of the waves, and sometimes by
their being completely blocked. Other methods by which the passage of
waves in contractile strips admits of being blocked will be alluded
to farther on.
Public-domain text, read in full here on John Shaqi.
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