The last-mentioned fact shows that after excitational continuity
of a higher order has been destroyed, excitational continuity of a
lower order nevertheless persists; or, to state the case in other
words, the fact in question shows that after severance of the almost
exclusively radial connections between the bell and the manubrium, by
which the perfect or unimpaired localizing function of the latter
is rendered possible, other connections between these organs remain
which are not in any wise radial. I therefore next tested the degree
in which these non-radial connections might be cut without causing
destruction of that excitational continuity of a lower order which
it is their function to maintain. It will here suffice to record one
mode of section which has yielded definite results. A glance at the
accompanying illustration (Fig. 23) will show the manner in which
the Medusa is prepared. The margin having been removed (in order to
prevent possible conduction by the marginal nerve-fibres), a single
deep radial cut (_a_ _a_) is first made, and then a circumferential
cut (_a_, _b_, _c_) is carried nearly all the way round the base of
the manubrium. In this way the nectocalyx, deprived of its margin,
is converted into a continuous band of tissue, one of the ends of
which supports the manubrium. Now it is obvious that this mode of
section must be very trying to nervous connections of any kind
subsisting between the bell and the manubrium. Nevertheless, in many
cases, irritating any part of the band _a_ _l_ has the effect of
causing the manubrium to perform the active random motions previously
described. In such cases, however, it is observable that the further
away from the manubrium the stimulus is applied, the less active
is the response of this organ. In very many instances, indeed, the
manubrium altogether fails to respond to stimuli applied at more than
a certain distance from itself. For example, referring to Fig. 23,
the manubrium might actively respond to irritation of any point in
the division _d_, _e_, _f_, _g_, while to irritation of any point
in the division _f_, _g_, _h_, _i_ its responses would be weaker,
and to irritation of any point in _h_, _i_, _j_, _k_, they would be
very uncertain or altogether absent. Hence in this form of section
we have reached about the limit of tolerance of which the non-radial
connections between the bell and manubrium are capable.
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Elsewhere in the archive
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account