Noticeable currents are not always produced when the mouth of this
Floscule is fully expanded. On one occasion, one having five lobes was
discovered standing at such an angle in a glass trough that the aperture
could be looked down into. The position rendered it impossible to use a
higher power than about two hundred linear, but with this, and the
employment of carmine, nothing like a vortex was seen during a whole
evening, although a less power was sufficient to show the ciliary
whirlpools made by small specimens of _Epistylis_ and _Vaginicola_,
which were in the small vessel. The density of the integument was
unfavorable to viewing the action of the gizzard, but it could be
indistinctly perceived. The contractions and subsequent expansions of
the cup, formed by the upper part of the creature, may be one way in
which its food is drawn in, but there is no doubt it can produce
currents when it thinks proper. Sometimes animalcules in the vicinity of
Floscules whirl about as if under the influence of such currents. Some
may be seen to enter the space between the lobes, swim about inside, and
then get out again, while every now and then one will be sucked in too
far for retreat.
Above the gizzard in the Horned Floscule,[10] I have seen an appearance
as if a membrane or curtain was waving to and fro, while another was
kept in a fixed perpendicular position. Mr. Gosse, speaking of this
genus, observes "that the whole of the upper part of the body is lined
with a sensitive, contractile, partially opaque membrane, which a little
below the disk recedes from the walls of the body, and forms a
diaphragm, with a highly contractile and versatile central orifice. At
some distance lower down another diaphragm occurs, and the ample chamber
thus enclosed forms a kind of _crop_, or receptacle for the captured
prey."
[10] The Horned Floscules (_F. cornuta_) which I have found, and which
bred in a glass jar, were not so large as those described by Mr. Dobie,
as quoted in 'Pritchard's Infusoria.' Mr. Dobie's specimens were 1--40"
when extended; mine about half that size, five-lobed, and with a long
slender proboscis, standing in a wavy line outside one lobe. Mr. Dobie
also describes an _F. campanulata_, with five flattened lobes. The
'Micrographic Dictionary' pronounces these two species "doubtfully
distinct." I have three or four times met with a variety of _F. ornata_,
in which one lobe was much enlarged and flattened, but they had no
proboscis. In what I take for _F. cornuta_, the horn or proboscis has
sometimes been a conspicuous object, and at others so fine and
transparent as to be only visible in certain lights.
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