Another kind of rotifer was abundant--the _Philodina_, which belongs to
the same family as the common wheel-bearer, namely, the _Philodinæa_.
The _Philodina_ is a good deal like the common wheel-bearer, or _Rotifer
vulgaris_, but is usually of a stouter build, and carries his eyes in a
different place. In the common rotifer these organs are situated on the
proboscis, while those of the Philodina are lower, and said to be
"cervical." The changes of form in this rotifer are still more
remarkable than in the common wheel-bearer. When resting it resembles a
pear-shaped purse, puckered in at the mouth. Then it thrusts out its
tail-foot, swells its body to an oval globe, protrudes its feeler, and
slightly exposes a row of cilia. After this two distinct wheels are
everted, and as their cilia whirl and spin, the animal is swiftly rowed
along, until it thinks proper to moor itself fast by the tail-foot, and
employ all its ciliary power in causing currents to converge towards its
throat. When it pleases it can elongate the body, till it becomes
vermiform, and it walks like the common rotifer, by curving its back,
and bringing its nose and its tail in contact with the ground.
[Illustration: Philodina (crawling).]
The gizzard of this family (_Philodinæa_) presents a considerable
deviation from the perfect form exhibited by the _Brachions_. According
to Mr. Gosse, "The _mallei_ and the _incus_ (terms already explained)
are soldered together into two subquadrantic-globular masses, which
appear to be muscular, but invested with a solid integument. The
_manubria_ (handles) may still be recognised in a vertical aspect as
three loops, of which the central one is chiefly developed, and in a
vertical aspect as a translucent reniform (kidney-shaped) globe." These
descriptions are not easy to understand, not from any want of clearness
or precision in the words employed, but from the complicated character
of the organ, and its very different appearance under different aspects.
To make the matter more intelligible, Mr. Gosse adds, "the structure and
action of an apparatus of this type may be made more clear by a homely
illustration. Suppose an apple to be divided longitudinally, leaving the
stalk attached to one half. Let this now be split again longitudinally
so far as the stalk, but not actually separating either portion from it.
Draw the two portions slightly apart, and lay them down on their rounded
surfaces. They now represent the quadrantic masses in repose, the stalk
being the fulcrum, and the upper surfaces being crossed by the teeth. By
the contraction of the muscles, of which they are composed, the two
segments are made to turn upon their long axis, until the points of the
teeth are brought into contact, and the toothed surfaces rise and
approach each other. The lower edges do not, however, separate as the
upper edges approach, but the form of the mass alters, becoming more
lenticular, so that when the toothed surfaces are brought into their
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