Mendel's principles of heredity: A defenceMendel, Gregor
Science
Mendel's principles of heredity: A defence
Mendel, Gregor
Heredity; Hybridization; Mendel's law; Weldon, Walter Frank Raphael. Mendel's laws of alternative inheritance in peas
2. _Datura._ As for the evidence on _Datura_, I must refer the reader
again to the experiments set forth in our Report.
The phenomena obey the ordinary Mendelian rules with accuracy. There
are (as almost always where discontinuous variation is concerned)
occasional cases of “mosaics,” a phenomenon which has nothing to do
with “ancestry.”
3. _Colours of Rats and Mice._ Professor Weldon reserves his
collection of evidence on this subject for the last. In it we reach an
indisputable contribution to the discussion--a reference to Crampe’s
papers, which together constitute without doubt the best evidence yet
published, respecting colour-heredity in an animal. So far as I have
discovered, the only previous reference to these memoirs is that of
Ritzema Bos[133], who alludes to them in a consideration of the alleged
deterioration due to in-breeding.
[133] _Biol. Cblt._ XIV. 1894, p. 79.
Now Crampe through a long period of years made an exhaustive study of
the peculiarities of the colour-forms of Rats, white, black, grey and
their piebalds, as exhibited in Heredity.
Till the appearance of Professor Weldon’s article Crampe’s work was
unknown to me, and all students of Heredity owe him a debt for putting
it into general circulation. My attention had however been called
by Dr Correns to the interesting results obtained by von Guaita,
experimenting with crosses originally made between albino _mice_ and
piebald Japanese waltzing mice. This paper also gives full details of
an elaborate investigation admirably carried out and recorded.
In the light of modern knowledge both these two researches furnish
material of the most convincing character demonstrating the Mendelian
principles. It would be a useful task to go over the evidence they
contain and rearrange it in illustration of the laws now perceived. To
do this here is manifestly impossible, and it must suffice to point
out that the albino is a simple recessive in both cases (the waltzing
character in mice being also a recessive), and that the “wild grey”
form is one of the commonest heterozygotes--there appearing, like
the yellow cotyledon-colour of peas, _in either of two capacities_,
i.e. as a pure form, or as the heterozygote form of one or more
combinations[134].
[134] The various “contradictions” which Professor Weldon suggests
exist between Crampe, von Guaita and Colladon can almost certainly be
explained by this circumstance. For Professor Weldon “wild-coloured”
mice, however produced, are “wild-coloured” mice and no more (see
Introduction).
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