Mendel's principles of heredity: A defenceMendel, Gregor
Science
Mendel's principles of heredity: A defence
Mendel, Gregor
Heredity; Hybridization; Mendel's law; Weldon, Walter Frank Raphael. Mendel's laws of alternative inheritance in peas
That we are in the presence of a new principle of the highest
importance is manifest. To what further conclusions it may lead us
cannot yet be foretold. But both Mendel and the authors who have
followed him lay stress on one conclusion, which will at once suggest
itself to anyone who reflects on the facts. For it will be seen that
the results are such as we might expect if it be imagined that the
cross-bred plant produced pollen grains and egg-cells, each of which
bears only _one_ of the alternative varietal characters and not both.
If this were so, and if on an average the same number of pollen grains
and egg-cells transmit each of the two characters, it is clear that on
a random assortment of pollen grains and egg-cells Mendel’s law would
be obeyed. For 25 per cent. of “dominant” pollen grains would unite
with 25 per cent. “dominant” egg-cells; 25 per cent. “recessive” pollen
grains would similarly unite with 25 per cent. “recessive” egg-cells;
while the remaining 50 per cent. of each kind would unite together.
It is this consideration which leads both Mendel and those who have
followed him to assert that these facts of crossing prove that each
egg-cell and each pollen grain is pure in respect of each character
to which the law applies. It is highly desirable that varieties
differing in the form of their pollen should be made the subject of
these experiments, for it is quite possible that in such a case strong
confirmation of this deduction might be obtained. [Preliminary trials
made with reference to this point have so far given negative results.
Remembering that a pollen grain is not a germ-cell, but only a bearer
of a germ-cell, the hope of seeing pollen grains differentiated
according to the characters they bear is probably remote. Better hopes
may perhaps be entertained in regard to spermatozoa, or possibly female
cells.]
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