Mendel's principles of heredity: A defenceMendel, Gregor
Science
Mendel's principles of heredity: A defence
Mendel, Gregor
Heredity; Hybridization; Mendel's law; Weldon, Walter Frank Raphael. Mendel's laws of alternative inheritance in peas
4. To the _difference in the form of the ripe pods_. These are
either simply inflated, never contracted in places; or they are
deeply constricted between the seeds and more or less wrinkled (_P.
saccharatum_).
5. To the _difference in the colour of the unripe pods_. They are
either light to dark green, or vividly yellow, in which colouring the
stalks, leaf-veins, and calyx participate[26].
[26] One species possesses a beautifully brownish-red coloured pod,
which when ripening turns to violet and blue. Trials with this
character were only begun last year. [Of these further experiments it
seems no account was published. Correns has since worked with such a
variety.]
6. To the _difference in the position of the flowers_. They are either
axial, that is, distributed along the main stem; or they are terminal,
that is, bunched at the top of the stem and arranged almost in a false
umbel; in this case the upper part of the stem is more or less widened
in section (_P. umbellatum_)[27].
[27] [This is often called the Mummy Pea. It shows slight fasciation.
The form I know has white standard and salmon-red wings.]
7. To the _difference in the length of the stem_. The length of the
stem[28] is very various in some forms; it is, however, a constant
character for each, in so far that healthy plants, grown in the same
soil, are only subject to unimportant variations in this character.
[28] [In my account of these experiments (_R.H.S. Journal_, vol. XXV.
p. 54) I misunderstood this paragraph and took “axis” to mean the
_floral_ axis, instead of the main axis of the plant. The unit
of measurement, being indicated in the original by a dash (′), I
carelessly took to have been an _inch_, but the translation here
given is evidently correct.]
In experiments with this character, in order to be able to discriminate
with certainty, the long axis of 6–7 ft. was always crossed with the
short one of 3/4 ft. to 1-1/2 ft.
Each two of the differentiating characters enumerated above were united
by cross-fertilisation. There were made for the
1st trial 60 fertilisations on 15 plants.
2nd " 58 " " 10 "
3rd " 35 " " 10 "
4th " 40 " " 10 "
5th " 23 " " 5 "
6th " 34 " " 10 "
7th " 37 " " 10 "
From a larger number of plants of the same variety only the most
vigorous were chosen for fertilisation. Weakly plants always afford
uncertain results, because even in the first generation of hybrids,
and still more so in the subsequent ones, many of the offspring either
entirely fail to flower or only form a few and inferior seeds.
Furthermore, in all the experiments reciprocal crossings were effected
in such a way that each of the two varieties which in one set of
fertilisations served as seed-bearers in the other set were used as
pollen plants.
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