Mendel's principles of heredity: A defenceMendel, Gregor
Science
Mendel's principles of heredity: A defence
Mendel, Gregor
Heredity; Hybridization; Mendel's law; Weldon, Walter Frank Raphael. Mendel's laws of alternative inheritance in peas
The plants were grown in garden beds, a few also in pots, and were
maintained in their naturally upright position by means of sticks,
branches of trees, and strings stretched between. For each experiment
a number of pot plants were placed during the blooming period in a
greenhouse, to serve as control plants for the main experiment in the
open as regards possible disturbance by insects. Among the insects[29]
which visit Peas the beetle _Bruchus pisi_ might be detrimental to the
experiments should it appear in numbers. The female of this species is
known to lay the eggs in the flower, and in so doing opens the keel;
upon the tarsi of one specimen, which was caught in a flower, some
pollen grains could clearly be seen under a lens. Mention must also be
made of a circumstance which possibly might lead to the introduction
of foreign pollen. It occurs, for instance, in some rare cases that
certain parts of an otherwise quite normally developed flower wither,
resulting in a partial exposure of the fertilising organs. A defective
development of the keel has also been observed, owing to which the
stigma and anthers remained partially uncovered[30]. It also sometimes
happens that the pollen does not reach full perfection. In this event
there occurs a gradual lengthening of the pistil during the blooming
period, until the stigmatic tip protrudes at the point of the keel.
This remarkable appearance has also been observed in hybrids of
_Phaseolus_ and _Lathyrus_.
[29] [It is somewhat surprising that no mention is made of Thrips,
which swarm in Pea flowers. I had come to the conclusion that this is
a real source of error and I see Laxton held the same opinion.]
[30] [This also happens in Sweet Peas.]
The risk of false impregnation by foreign pollen is, however, a very
slight one with _Pisum_, and is quite incapable of disturbing the
general result. Among more than 10,000 plants which were carefully
examined there were only a very few cases where an indubitable false
impregnation had occurred. Since in the greenhouse such a case was
never remarked, it may well be supposed that _Bruchus pisi_, and
possibly also the described abnormalities in the floral structure, were
to blame.
THE FORMS OF THE HYBRIDS.[31]
[31] [Mendel throughout speaks of his cross-bred Peas as “hybrids,” a
term which many restrict to the offspring of two distinct _species_.
He, as he explains, held this to be only a question of degree.]
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