Though the Pharmacophagus Papilios are mimicked only by other Papilios
among butterflies they may serve occasionally as models for certain of the
larger day-flying moths. _Papilio polyxenus_, for example, is mimicked not
only by the unprotected _P. bootes_ but also by the moth _Epicopeia
polydora_ (Pl. III, figs. 5 and 6). Like the butterfly the _Epicopeia_,
which is comparatively rare, has the white patch and the outer border of
red marginal spots on the hind wing. Though it is apparently unable to
provide itself with an orthodox tail it nevertheless makes a creditable
attempt at one. There are several other cases of mimetic resemblance
between day-flying moths and Pharmacophagus swallow-tails--the latter in
each case serving as the model. Rarely it may happen that the rôle of
butterfly and moth is reversed, and the butterfly becomes the mimic. A very
remarkable instance of this is found in New Guinea where the rare _Papilio
laglaizei_ mimics the common day-flying moth _Alcidis agathyrsus_. Viewed
from above the resemblance is sufficiently striking (Pl. III, figs. 1 and
2), but the most wonderful feature concerns the underneath. The ventral
half of the moth's abdomen is coloured brilliant orange. When the wings are
folded back they cover and hide from sight only the dorsal part of the
abdomen, so that in this position the orange neutral surface is
conspicuous. When, however, the wings of the butterfly are folded they
conceal the whole of the abdomen. But the butterfly has developed on each
hind wing itself a bright orange patch in such a position that when the
{28} wings are folded back the orange patch lies over the sides of the
abdomen. In this way is simulated the brilliant abdomen of the moth by a
butterfly, in which, as in its relations, this part is of a dark and sombre
hue.
A few models are also provided in the Oriental region by the genus
_Delias_, which belongs to the Pierines. A common form, _Delias eucharis_,
is white above but the under surface of the hind wings is conspicuous with
yellow and scarlet (Pl. II, fig. 1). It has been suggested that this
species serves as a model for another and closely allied Pierine,
_Prioneris sita_, a species distinctly scarcer than the _Delias_. There is
some evidence that the latter is distasteful (cf. p. 115), but nothing is
known of the _Prioneris_ in this respect. Other species of _Delias_ are
said to function as models for certain day-flying moths belonging to the
family Chalcosiidae, which may bear a close resemblance to them. In certain
cases it may happen that the moth is more abundant than the Pierine that it
resembles[12].
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