================================================================== |Snout-vent |Head width/|Interorbital Locality |length |snout-vent |distance/ | |length |head width -----------------------------+-----------+-----------+------------ Bonanza, Nicaragua | 40.8-47.7 | 34.1-38.0 | 31.0-36.1 | (43.7) | (36.3) | (35.4) -----------------------------+-----------+-----------+------------ Heredia Prov., Costa Rica | 46.3-59.0 | 32.5-36.0 | 30.5-39.6 | (51.7) | (35.0) | (34.7) -----------------------------+-----------+-----------+------------ Puntarenas Prov., Costa Rica | 53.6-64.9 | 32.6-36.1 | 31.0-38.0 | (61.4) | (34.5) | (34.4) -----------------------------+-----------+-----------+------------ Canal Zone, Panama | 52.4-65.5 | 33.5-37.6 | 31.3-37.2 | (56.5) | (35.6) | (34.7) -----------------------------+-----------+-----------+------------ Rio Quesada, Colombia | 52.6-61.0 | 33.1-37.1 | 30.1-33.9 | (56.0) | (35.0) | (32.1) ------------------------------------------------------------------ _Description and Variation._--For the purposes of analyzing geographic variation in size and proportions, measurements were taken on ten adult males from each of five samples throughout the range of the species. Aside from the data summarized in Table 2, the average ratio of tibia length to snout-vent length is noticeably less in Colombian specimens (53.4 per cent, as compared with 54.8 to 57.8 per cent in the other samples) and the ratio of head length to snout-vent length is noticeably less in Costa Rican specimens (33.5 per cent as compared with 34.9 to 35.1 per cent in the other samples). Also, specimens from Heredia Province, Costa Rica, have a relatively smaller tympanum (62.7 to 80.4 [mean 68.4] per cent of the diameter of the eye, as compared with means of 74.0 to 77.9 per cent in the other samples). Two populations are distinctive as regards the size of adult males. Specimens from the northern Caribbean lowlands of Nicaragua (Bonanza, the northernmost locality for the species) are remarkably small. Males having snout-vent lengths of between 40 and 43 mm. were breeding; the largest male found had a snout-vent length of 47.7 mm. The other extreme in size is attained in specimens from the Pacific lowlands of eastern Costa Rica and western Panama, where most breeding males have snout-vent lengths of more than 55 mm.; the largest male had a snout-vent length of 64.9 mm.