North American Recent Soft-Shelled Turtles (Family Trionychidae) — John Shaqi
North American Recent Soft-Shelled Turtles (Family Trionychidae)Webb, Robert G. (Robert Gravem)
Science
North American Recent Soft-Shelled Turtles (Family Trionychidae)
Webb, Robert G. (Robert Gravem)
Soft-shelled turtles
_Synonomy._--Geoffroy published a synopsis of the species he
recognized (1809) prior to his formal description of the genus
_Trionyx_ (1809a). Schweigger, nevertheless, probably was the first
person to recognize the soft-shelled turtles as a distinct group, and
he proposed for it the name _Amyda_ in an unpublished manuscript that
he sent to Geoffroy. The latter author (1809a:15) relegated the name
_Amyda_ to the synonomy of _Trionyx javanicus_ by means of the
following entry: "_Amyda javanica._ Schweigger, dans un manuscript
communique a l'Institut." Stejneger (1944:7) maintained that this
publication of Schweigger's monotypic generic name clearly established
its availability for the species congeneric with _Amyda javanica_ (=
_Testudo cartilaginea_ Boddaert, 1770). Loveridge and Williams
(1957:422) contend that this mere mention of the name _Amyda_ neither
constitutes the proposal of a new name nor validates it, and that the
first valid usage of the name _Amyda_ is that of Fitzinger (1835:120),
who later (1843:30) designated the type species as _Amyda subplana_.
The name _Amyda_ cannot date from _Oken_ (1816:348) as Volume 3
[Zoologie] of his Lehrbuch der Naturgeschichte published in 1815-1816
has been placed on the Official Index of Rejected and Invalid Works in
Zoological Nomenclature with the Title No. 33; see Opinion 417
(Hemming, 1956).
There has been considerable debate as to whether Geoffroy did or did
not designate a type species of the genus _Trionyx_ (1809a). Although
not specifically designated as the type species, _Trionyx aegyptiacus_
(= _Testudo triunguis_ Forskål) is considered by Smith (1930:2),
Schmidt (1953:108, footnote), and Loveridge and Williams (1957:422) to
have been sufficiently indicated by Geoffroy as the type species. But
Stejneger (1944:6), H. M. Smith (1947:122), Conant and Goin
(1948:11), and Mertens and Wermuth (1955) maintained that Geoffroy did
not adequately designate a type species, and that Fitzinger (1843:30)
designated the type species as _Trionyx granosus_ (= _Lissemys
punctata_), a synonym of Geoffroy's species, _coromandelicus_.
If Fitzinger's designation of a type species is accepted, the name
_Trionyx_ is applicable to the forms herein referred to _Lissemys_,
and _Amyda_ to the American forms. If Geoffroy's designation is
accepted, the American forms are referable to _Trionyx_, and _Amyda_
is a synonym.
The preceding includes only those generic names (listed in
chronological order) that have been applied to Recent American
soft-shelled turtles. Generic synonyms of the genus _Trionyx_
applicable to Old World species are listed by Stejneger (1907:514),
Smith (1931:165), and Loveridge and Williams (1957:420-21).
_Trionyx_ is the most widespread genus of the family; most of the
species occur in southeastern Asia. All North American soft-shelled
turtles belong to this genus.
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