Observations on the Terataspis grandis, Hall, the largest known trilobite — John Shaqi
Observations on the Terataspis grandis, Hall, the largest known trilobiteClarke, John Mason
Science
Observations on the Terataspis grandis, Hall, the largest known trilobite
Clarke, John Mason
Paleontology -- New York (State); Trilobites
Not until the introduction of the genera DALMANITES, HOMALONOTUS
and LICHAS do we meet with the most gigantic proportions attained
by these crustaceans, and then only after these genera have become
well established. Perhaps none of their representatives in the lower
Silurian faunas were of commanding size; in the upper Silurian large
but not extravagant proportions were sometimes attained. _Lichas
Boltoni_ of the Niagara fauna, is a magnificent species, one of the
largest of its race and remarkable for the frequency with which its
parts are found together, an extremely uncommon occurrence in this
thin-shelled group. The _Lichas pustulosus_ of the Lower Helderberg
shaly limestone was a great species attaining a length of 10 inches
or more. _Homalonotus delphinocephalus_ of the Niagara fauna grew to
large size but does not appear to have attained the length of its
successor in the Lower Helderberg, _H. Vanuxemi_, which, according to
the restoration from a very large fragment given in the Palæontology
of New York, Volume VII (plate V B) grew to a length of at least 11½
inches. SALTER has mentioned (Palæontographical Society, vol. xvii, p.
109) a large fragment of _Homalonotus rudis_ which he estimates may
have been a foot in length. In later faunas are found traces of this
genus of still greater size. Dr. BRUSHAUSEN has figured a pygidium of
_H. gigas_ from the Spiriferen-sandstein of the Hartz, the possessor
of which must have been upward of one foot in length. Unquestionably
the largest individual of HOMALONOTUS known is that of _H. major_,
from the Oriskany sandstone, figured in the Palæontology of New York,
Volume VII (plate V A), a large fragment representing the greater part
of the thorax and the pygidium, and according to the restoration there
given the original length of the animal must have been well nigh 15
inches. The later and common Hamilton species, _H. DeKayi_, attained no
extravagant size though frequently large, e. g. the enrolled individual
figured on plate IV (_op. cit._), the largest entire specimen reported,
which is about 9 inches in length; some large fragments indicate that
the animal was sometimes as long as 11 inches.
The size attained by some of the Devonian species of DALMANITES and
their immediate predecessors was marvelous. The pygidium of _D.
micrurus_ figured in the Palæontology of New York, Volume III, page
359 (there given as _D. pleuroptyx_), indicates an individual at least
11 inches in length, and there is reason to believe that the Lower
Helderberg species, _D. nasutus_ and _D. tridens_, attained a size
fully as great. Most remarkable however is the great pygidium of _D.
myrmecophorus_ of the Corniferous limestone, figured upon plate XV of
Volume VII (_op. cit._), which from the restoration there given, made
from careful comparative measurements, would imply an individual 16
inches long.
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