On Germinal Selection as a Source of Definite Variation — John Shaqi
On Germinal Selection as a Source of Definite VariationWeismann, August
Science
On Germinal Selection as a Source of Definite Variation
Weismann, August
Variation (Biology)
Discouraging, therefore, as it may be, that the control of nature in her
minutest details is here gainsaid us, yet it were equivalent to sacrificing
the gold to the dross, if simply from our inability to follow out the
details of the individual case we should renounce altogether the principle
of selection, or should proclaim it as only subsidiary, on the ground that
we believe the protective coloring of the butterfly is not a protective
coloring, but a combination of colors inevitably resulting from internal
causes. The protective coloring remains a protective coloring whether at
the time in question it is or is not necessary for the species; and it
arose as protective coloring--arose not because it was a constitutional
necessity of the animal's organism that here a red and there a white,
black, or yellow spot should be produced, but because it was {16}
advantageous, because it was necessary for the animal. There is only one
explanation possible for such patent adaptations and that is selection.
What is more, no other natural way of their originating is conceivable, for
we have no right to assume teleological forces in the domain of natural
phenomena.
I have selected the example of the butterfly's wing, not solely because it
is so widely known, but because it is so exceedingly instructive, because
we are still able to learn so much from it. It has been frequently asserted
that the color-patterns of the butterfly's wings have originated from
internal causes, independently of selection and conformably to inward laws
of evolution. Eimer has attempted to prove this assertion by establishing
in a division of the genus Papilio the fact that the species there admit of
arrangement in series according to affinity of design. But is a proof that
the markings are modified in definite directions during the course of the
species's development equivalent to a definite statement as to the _causes_
that have produced these gradual transformations? Or, is our present
inability to determine with exactness the biological significance of these
markings and their modifications, a proof that the same have no
significance whatever? On the contrary, I believe it can be clearly proved
that the wing of the butterfly is a tablet on which nature has inscribed
everything she has deemed advantageous to the preservation and welfare of
her creatures, and nothing else; or, to abandon the simile, that these
color-patterns have not proceeded from inward evolutional forces, but are
the result of selection. At least in all places where we do understand
their biological significance these patterns are constituted and
distributed over the wing exactly as utility would require. {17}
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