On Germinal Selection as a Source of Definite VariationWeismann, August
Science
On Germinal Selection as a Source of Definite Variation
Weismann, August
Variation (Biology)
Should any one be still unconvinced by the evidence we have adduced, let
him give the leaf-markings a closer inspection. He will find that the
midrib is composed of two pieces of which the one belongs to the {24} hind
wing and the other to the fore wing, and that the two fit each other
exactly when the butterfly is in the attitude of repose, but not otherwise.
Now these two pieces of the leaf-rib do not begin on corresponding spots of
the two wings, but on absolutely non-identical spots. And the same is also
true of the lines which represent the lateral ribs of the leaf. These lines
proceed in acute angles from the rib; to the right and to the left in the
same angle, those of the same side parallel with each other. Here, too, no
relation is noticeable between the parts of the wings over which the lines
pass. The venation of the wing is utterly ignored by the leaf-markings, and
its surface is treated as a _tabula rasa_ upon which anything conceivable
can be drawn. In other words, we are presented here with a _bilaterally
symmetrical_ figure engraved on a surface which is essentially _radially
symmetrical_ in its divisions.
I lay unusual stress upon this point because it shows that we are dealing
here with one of those cases which cannot be explained by mechanical, that
is, by natural means, unless natural selection actually exists and is
actually competent to create new properties; for the Lamarckian principle
is excluded here _ab initio_, seeing that we are dealing with a formation
which is only passive in its effects; the leaf-markings are effectual
simply by their existence and not by any function which they perform; they
are present in flight as well as at rest, during the absence of danger, as
well as during the approach of an enemy.
Nor are we helped here by the assumption of _purely internal motive
forces_, which Naegeli, Askenasy, and others have put forward as supplying
a _mechanical_ force of evolution. It is impossible to regard the {25}
coincidence of an Indian butterfly with the leaf of a tree now growing in
an Indian forest as fortuitous, as a _lusus naturae_. Assuming this
seemingly mechanical force, therefore, we should be led back inevitably to
a teleological principle which produces adaptive characters and which must
have deposited the directive principle in the very first germ of
terrestrial organisms, so that after untold ages at a definite time and
place the illusive leaf-markings should be developed. The assumption of
pre-established harmony between the evolution of the ancestral line of the
tree with its pre-figurative leaf, and that of the butterfly with its
imitating wing, is absolutely necessary here--a fact which I pointed out
many years ago,[9] but which is constantly forgotten by the promulgators of
the theory of internal evolutionary forces.
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account