On Germinal Selection as a Source of Definite Variation — John Shaqi
On Germinal Selection as a Source of Definite VariationWeismann, August
Science
On Germinal Selection as a Source of Definite Variation
Weismann, August
Variation (Biology)
The Lamarckians were right when they maintained that the factor for which
hitherto the name of natural selection had been exclusively reserved, viz.,
_personal_ selection, was insufficient for the explanation of the
phenomena. They were also right when they declared that panmixia in the
form in which until recently I held the theory was also insufficient to
explain the degeneration of parts that had grown useless, but they {65}
erred when they ascribed hereditary effects to the selection-processes
which are enacted among the parts of the body (Wilhelm Roux) and which are
rightly regarded as the results of functioning. And they did this,
moreover, as they themselves admit, not because the facts of heredity
directly and unmistakably required it, but because they saw no other
possibility of explaining many phenomena of transformation. I am fain to
relinquish myself to the hope that now after another explanation has been
found, a reconciliation and unification of the hostile views is not so very
distant, and that then, we can continue our work together on the newly laid
foundations.
That the application of the Malthusian principle was thoroughly justified
is now clear. _The entire process of the development of living forms is
guided by this principle._ The struggle for existence, _videlicet_, for
food and propagation, takes place at all the stages of life between all
orders of living units from the biophores recently disclosed upwards to the
elements that are accessible to direct observation, to the cells, and still
higher up, to individuals and colonies. Consequently, in all the divers
orders of biological units lying between the two extremes of biophores and
colonies, the modifications must be controlled by selective processes;
therefore, these govern every change of living forms no matter what its
significance, and bring it about that the latter fit their conditions of
life as wax does the mould; and the various stages of these processes, as
enacted between the divers orders of biological units, in all organisms not
absolutely simple, are involved in incessant and mutual interaction. The
three principal stages of selection, that of {66} _personal_ selection[23]
as it was enunciated by Darwin and Wallace, that of _histonal_ selection as
it was established by Wilhelm Roux in the form of a "struggle of the
parts," and finally that of _germinal selection_ whose existence and
efficacy I have endeavored to substantiate in this article--these are the
factors that have co-operated to maintain the forms of life in a constant
state of viability and to adapt them to their conditions of life, now
modifying them _pari passu_ with their environment, and now maintaining
them on the stage attained, when that environment is not altered.
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