On Germinal Selection as a Source of Definite Variation — John Shaqi
On Germinal Selection as a Source of Definite VariationWeismann, August
Science
On Germinal Selection as a Source of Definite Variation
Weismann, August
Variation (Biology)
Everything is adapted in animate nature[24] and has been from the first
beginnings of life; for adaptiveness of organisation is here equivalent to
the power to exist, and they alone have had the power to exist who have
permanently existed. _We know of only one natural principle of explanation
for this fact--that of selection {67} of the picking out of those having
the power to exist from those having the power to originate._ If there is
any solution possible to the riddle of adaptiveness to ends,--a riddle held
by former generations to be insoluble,--it can be obtained only through the
assistance of this principle of the self-regulation of the originating
organisms, and we should not turn our faces and flee at the sight of the
first difficulties that meet its application, but should look to it whether
the apparent effects of this single principle of explanation are not
founded in the imperfect application that is made of it.
If I am not mistaken the situation is as follows: We had remained standing
half way. We had applied the principle, but only to a portion of the
natural units engaged in struggle. If we apply the principle throughout we
reach a satisfactory explanation. Selection of _persons_ alone is _not
sufficient_ to explain the phenomena; _germinal_ selection must be added.
Germinal selection is the last consequence of the application of the
principle of Malthus to living nature. It is true it leads us into a
terrain which cannot be submitted directly to observation by means of our
organs of touch and by our eyes, but it shares this disadvantage in common
with all other ultimate inferences in natural science, even in the domain
of inorganic {68} nature: in the end all of them lead us into hypothetical
regions. If we are not disposed to follow here, nothing remains but to
abandon utterly the hope of explaining the adaptive character of life--a
renunciation which is not likely to gain our approval when we reflect that
by the other method is actually offered at least in principle, not only a
broad insight into the adaptation of the single forms of life to their
conditions, but also into the mode of formation of the living world as a
whole. The variety of the organised world, its transformation by adaptation
to new, and by reversed adaptation to old conditions, the inequality of the
systematic groups, the attainment of the same ends by different means, that
is, by different organisations, and a thousand and one other things assume
on this hypothesis in a certain measure an intelligible form, whilst
without it they remain lifeless facts.
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