On Germinal Selection as a Source of Definite VariationWeismann, August
Science
On Germinal Selection as a Source of Definite Variation
Weismann, August
Variation (Biology)
FREIBURG, Nov. 18, 1895.
* * * * *
{11}
GERMINAL SELECTION.
* * * * *
Numerous and varied are the objections that have been advanced against the
theory of selection since it was first enunciated by Darwin and
Wallace--from the unreasoning strictures of Richard Owen and the acute and
thoughtful criticisms of Albert Wigand and Naegeli to the opposition of our
own day, which contends that selection cannot create but only reject, and
which fails to see that precisely through this rejection its creative
efficacy is asserted. The champions of this view are for discovering the
motive forces of evolution in the _laws_ that govern organisms--as if the
norm according to which an event happens were the event itself, as if the
rails which determine the direction of a train could supplant the
locomotive. Of course, from every form of life there proceeds only a
definite, though extremely large, number of tracks, _the possible
variations_, whilst between them lie stretches without tracks, _the
impossible variations_, on which locomotion is impossible. But the actual
travelling of a track is not performed by the track, but by the locomotive,
and on the other hand, the choice of a track, the decision whether the
destination of the train shall be Berlin or Paris, is not made by the
locomotive, the cause of the variation, but by the driver of the
locomotive, who directs the engine on the right track. In the theory of
selection the engine-driver is represented by utility, for with utility
rests the decision {12} as to what particular variational track shall be
travelled. The cogency, the irresistible cogency, as I take it, of the
principle of selection is precisely its capacity of explaining why fit
structures always arise, and that certainly is the great problem of life.
Not the fact of change, but the _manner_ of the change, whereby all things
are maintained capable of life and existence, is the pressing question.
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