On the phenomena of variation and geographical distribution as illustrated by the Papilionidæ of the Malayan region — John Shaqi
On the phenomena of variation and geographical distribution as illustrated by the Papilionidæ of the Malayan regionWallace, Alfred Russel
Science
On the phenomena of variation and geographical distribution as illustrated by the Papilionidæ of the Malayan region
Wallace, Alfred Russel
Papilionidae -- Malaysia -- Geographical distribution; Papilionidae -- Malaysia -- Variation
But, besides the true _P. Polytes_, there are several allied forms of
females to be considered, namely, _P. Theseus_, Cr., _P. Melanides_, De
Haan, _P. Elyros_, G. R. G., and _P. Romulus_, L. The dark female
figured by Cramer as _P. Theseus_ seems to be the common and perhaps the
only form in Sumatra, whereas in Java, Borneo, and Timor, along with
males quite identical with those of Sumatra, occur females of the
_Polytes_ form, although a single specimen of the true _P. Theseus_,
Cr., taken at Lombock would seem to show that the two forms do occur
together. In the allied species found in the Philippine ♀Islands (_P.
Alphenor_, Cr., _P. Ledebouria_, Eschsch., ♀ _P. Elyros_, G. R. G.)
forms corresponding to these extremes occur along with a number of
intermediate varieties, as shown by a fine series in the British Museum.
We have here an indication of how dimorphism may be produced; for let
the extreme Philippine forms be better suited to their conditions of
existence than the intermediate connecting links, and the latter will
gradually die out, leaving two distinct forms of the same insect, each
adapted to some special conditions. As these conditions are sure to vary
in different districts, it will often happen, as in Sumatra and Java,
that the one form will predominate in the one island, the other in the
adjacent one. In the island of Borneo there seems to be a third form;
for _P. Melanides_, De Haan, evidently belongs to this group, and has
all the chief characteristics of _P. Theseus_, with a modified
coloration of the hind wings. I now come to an insect which, if I am
correct, offers one of the most interesting cases of variation yet
adduced. _Papilio Romulus_, L., a butterfly found over a large part of
India and Ceylon, and not uncommon in collections, has always been
considered a true and independent species, and no suspicions have been
expressed regarding it. But a male of this form does not, I believe,
exist. I have examined the fine series in the British Museum, in the
East India Company’s Museum, in the Hope Museum at Oxford, in Mr.
Hewitson’s and several other private collections, and can find nothing
but females; and for this common butterfly no male partner can be found
except the equally common _P. Pammon_, a species already provided with
two wives, and yet to whom we shall be forced, I believe, to assign a
third. On carefully examining _P. Romulus_, I find that in all essential
characters,—the form and texture of the wings, the length of the
antennæ, the spotting of the head and thorax, and even the peculiar
tints and shades with which it is ornamented,—it corresponds exactly
with the other females of the _Pammon_ group; and though, from the
peculiar marking of the fore wings, it has at first sight a very
different aspect, yet a closer examination shows that every one of its
markings could be produced by slight and almost imperceptible
modifications of the various allied forms. I fully believe, therefore,
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