On the phenomena of variation and geographical distribution as illustrated by the Papilionidæ of the Malayan regionWallace, Alfred Russel
Science
On the phenomena of variation and geographical distribution as illustrated by the Papilionidæ of the Malayan region
Wallace, Alfred Russel
Papilionidae -- Malaysia -- Geographical distribution; Papilionidae -- Malaysia -- Variation
that I shall be correct in placing _P. Romulus_ as a third Indian form
of the female _P. Pammon_, corresponding to _P. Melanides_, the third
form of the Malayan _P. Theseus_. I may mention here that the females of
this group have a superficial resemblance to the _Polydorus_ group, as
shown by _P. Theseus_ having been considered to be the female of _P.
Antiphus_, and by _P. Romulus_ being arranged next to _P. Hector_. There
is no close affinity between these two groups of _Papilio_, and I am
disposed to believe that we have here a case of mimicry, brought about
by the same causes which Mr. Bates has so well explained in his account
of Heliconidæ, and which thus led to the singular exuberance of
polymorphic forms in this and allied groups of the genus _Papilio._ I
shall have to devote a section of my paper to the consideration of this
subject.
The third example of polymorphism I have to bring forward is _Papilio
Ormenus_, Guér., which is closely allied to the well-known _P.
Erechtheus_, Don., of Australia. The most common form of the female also
resembles that of _P. Erechtheus_; but a totally different-looking
insect was found by myself in the Aru Islands, and figured by Mr.
Hewitson under the name of _P. Onesimus_, which subsequent observation
has convinced me is a second form of the female of _P. Ormenus_.
Comparison of this with Boisduval’s description of _P. Amanga_, a
specimen of which from New Guinea is in the Paris Museum, shows the
latter to be a closely similar form; and two other specimens were
obtained by myself, one in the island of Goram and the other in Waigiou,
all evidently local modifications of the same form. In each of these
localities males and ordinary females of _P. Ormenus_ were also found.
So far there is no evidence that these light-coloured insects are not
females of a distinct species, the males of which have not been
discovered. But two facts have convinced me this is not the case. At
Dorey, in New Guinea, where males and ordinary females closely allied to
_P. Ormenus_ occur (but which seem to me worthy of being separated as a
distinct species), I found one of these light-coloured females closely
followed in her flight by three males, exactly in the same manner as
occurs (and, I believe, occurs only) with the sexes of the same species.
After watching them a considerable time, I captured the whole of them,
and became satisfied that I had discovered the true relations of this
anomalous form. The next year I had corroborative proof of the
correctness of this opinion by the discovery in the island of Batchian
of a new species allied to _P. Ormenus_, all the females of which,
either seen or captured by me, were of one form, and much more closely
resembling the abnormal light-coloured females of _P. Ormenus_ and _P.
Pandion_ than the ordinary specimens of that sex. Every naturalist will,
I think, agree that this is strongly confirmative of the supposition
that both forms of female are of one species; and when we consider,
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