On the phenomena of variation and geographical distribution as illustrated by the Papilionidæ of the Malayan region — John Shaqi
On the phenomena of variation and geographical distribution as illustrated by the Papilionidæ of the Malayan regionWallace, Alfred Russel
Science
On the phenomena of variation and geographical distribution as illustrated by the Papilionidæ of the Malayan region
Wallace, Alfred Russel
Papilionidae -- Malaysia -- Geographical distribution; Papilionidae -- Malaysia -- Variation
favourable to the one form, _there_ to the other; but we are by no means
to suppose that these conditions consist in climate alone. It is highly
probable that the existence of enemies, and of competing forms of life,
may be the main determining influences; and it is much to be wished that
such a competent observer as Mr. Walsh would endeavour to ascertain what
are the adverse causes which are most efficient in keeping down the
numbers of each of these contrasted forms.
Dimorphism of this kind in the animal kingdom does not seem to have any
direct relations to the reproductive powers, as Mr. Darwin has shown to
be the case in plants, nor does it appear to be very general. One other
case only is known to me in another family of my eastern Lepidoptera,
the _Pierulæ_; and but few occur in the Lepidoptera of other countries.
The spring and autumn broods of some European species differ very
remarkably; and this must be considered as a phenomenon of an analogous
though not of an identical nature[3]. _Araschnia prorsa_, of Central
Europe, is a striking example of this alternate or seasonal dimorphism.
Mr. Pascoe has pointed out two forms of the male sex in some species of
Coleoptera belonging to the family Anthribidæ, in seven species of the
two genera _Xenocerus_ and _Mecocerus_ (Proc. Ent. Soc. Lond., 1862, p.
71); and no less than six European Water-beetles, of the genus
_Dytiscus_, have females of two forms, the most common having the elytra
deeply sulcate, the rarer smooth as in the males. The three, and
sometimes four or more, forms under which many Hymenopterous insects
(especially Ants) occur must be considered as a related phenomenon,
though here each form is specialized to a distinct function in the
economy of the species. Among the higher animals, albinoism and melanism
may, as I have already stated, be considered as analogous facts; and I
met with one case of a bird, a species of Lory (_Eos fuscata_, Blyth),
clearly existing under two forms, since I obtained both sexes of each
from a single flock.
Footnote 3:
Among our nocturnal Lepidoptera, I am informed, many analogous cases
occur; and as the whole history of many of these has been investigated
by breeding successive generations from the egg, it is to be hoped
that some of our British Lepidopterists will give us a connected
account of all the abnormal phenomena which they present.
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