On the phenomena of variation and geographical distribution as illustrated by the Papilionidæ of the Malayan region — John Shaqi
On the phenomena of variation and geographical distribution as illustrated by the Papilionidæ of the Malayan regionWallace, Alfred Russel
Science
On the phenomena of variation and geographical distribution as illustrated by the Papilionidæ of the Malayan region
Wallace, Alfred Russel
Papilionidae -- Malaysia -- Geographical distribution; Papilionidae -- Malaysia -- Variation
4. _Coexisting variety._—This is a somewhat doubtful case. It is when a
slight but permanent and hereditary modification of form exists in
company with the parent or typical form, without presenting those
intermediate gradations which would constitute it a case of simple
variability. It is evidently only by direct evidence of the two forms
breeding separately that this can be distinguished from _dimorphism_.
The difficulty occurs in _Papilio Jason_, Esp., and _P. Evemon_, Bd.,
which inhabit the same localities, and are almost exactly alike in form,
size, and coloration, except that the latter always wants a very
conspicuous red spot on the under surface, which is found not only in
_P. Jason_, but in all the allied species. It is only by breeding the
two insects that it can be determined whether this is a case of a
coexisting variety or of _dimorphism_. In the former case, however, the
difference being constant and so very conspicuous and easily defined, I
see not how we could escape considering it as a distinct species. A true
case of coexisting forms would, I consider, be produced, if a slight
variety had become fixed as a local form, and afterwards been brought
into contact with the parent species with little or no intermixture of
the two; and such instances do very probably occur.
5. _Race, or subspecies._—These are local forms completely fixed and
isolated; and there is no possible test but individual opinion to
determine which of them shall be considered as species and which
varieties. If stability of form and “_the constant transmission of some
characteristic peculiarity of organization_” is the test of a species
(and I can find no other test that is more certain than individual
opinion), then every one of these fixed races, confined as they almost
always are to distinct and limited areas, must be regarded as a species;
and as such I have in most cases treated them. The various modifications
of _Papilio Ulysses_, _P. Peranthus_, _P. Codrus_, _P. Eurypilus_, _P.
Helenus_, &c., are excellent examples; for while some present great and
well-marked, others offer slight and inconspicuous differences, yet in
all cases these differences seem equally fixed and permanent. If,
therefore, we call some of these forms species, and others varieties, we
introduce a purely arbitrary distinction, and shall never be able to
decide where to draw the line. The races of _Papilio Ulysses_, L., for
example, vary in amount of modification from the scarcely differing New
Guinea form to those of Woodlark Island and New Caledonia, but all seem
equally constant; and as most of these had already been named and
described as species, I have added the New Guinea form under the name of
_P. Penelope_. We thus get a little group of Ulyssine Papilios, the
whole comprised within a very limited area, each one confined to a
separate portion of that area, and, though differing in various amounts,
each apparently constant. Few naturalists will doubt that all these may
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