On the phenomena of variation and geographical distribution as illustrated by the Papilionidæ of the Malayan regionWallace, Alfred Russel
Science
On the phenomena of variation and geographical distribution as illustrated by the Papilionidæ of the Malayan region
Wallace, Alfred Russel
Papilionidae -- Malaysia -- Geographical distribution; Papilionidae -- Malaysia -- Variation
The phenomena of _dimorphism_ and _polymorphism_ may be well
illustrated by supposing that a blue-eyed, flaxen-haired Saxon man had
two wives, one a black-haired, red-skinned Indian squaw, the other a
woolly-headed, sooty-skinned negress—and that instead of the children
being mulattoes of brown or dusky tints, mingling the separate
characteristics of their parents in varying degrees, all the boys
should be pure Saxon boys like their father, while the girls should
altogether resemble their mothers. This would be thought a
sufficiently wonderful fact; yet the phenomena here brought forward as
existing in the insect-world are still more extraordinary; for each
mother is capable not only of producing male offspring like the
father, and female like herself, but also of producing other females
exactly like her fellow-wife, and altogether differing from herself.
If an island could be stocked with a colony of human beings having
similar physiological idiosyncrasies with _Papilio Pammon_ or _Papilio
Ormenus_, we should see white men living with yellow, red, and black
women, and their offspring always reproducing the same types; so that
at the end of many generations the men would remain pure white, and
the women of the same well-marked races as at the commencement.
3. _Local form, or variety._—This is the first step in the transition
from variety to species. It occurs in species of wide range, when groups
of individuals have become partially isolated in several points of its
area of distribution, in each of which a characteristic form has become
segregated more or less completely. Such forms are very common in all
parts of the world, and have often been classed as varieties or species
alternately. I restrict the term to those cases where the difference of
the forms is very slight, or where the segregation is more or less
imperfect. The best example in the present group is _Papilio Agamemnon_,
L., a species which ranges over the greater part of tropical Asia, the
whole of the Malay archipelago, and a portion of the Australian and
Pacific regions. The modifications are principally of size and form,
and, though slight, are tolerably constant in each locality. The steps,
however, are so numerous and gradual that it would be impossible to
define many of them, though the extreme forms are sufficiently distinct.
_Papilio Sarpedon_, L., presents somewhat similar but less numerous
variations.
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