I find as a general rule that the constancy of species is in an inverse ratio to their range. Those which are confined to one or two islands are generally very constant. When they extend to many islands, considerable variability appears; and when they have an extensive range over a large part of the Archipelago, the amount of unstable variation is very large. These facts are explicable on Mr. Darwin’s principles. When a species exists over a wide area, it must have had, and probably still possesses, great powers of dispersion. Under the different conditions of existence in various portions of its area, different variations from the type would be selected, and, were they completely isolated, would soon become distinctly modified forms; but this process is checked by the dispersive powers of the whole species, which leads to the more or less frequent intermixture of the incipient varieties, which thus become irregular and unstable. Where, however, a species has a limited range, it indicates less active powers of dispersion, and the process of modification under changed conditions is less interfered with. The species will therefore exist under one or more permanent forms according as portions of it have been isolated at a more or less remote period. What is commonly called variation consists of several distinct phenomena which have been too often confounded. I shall proceed to consider these under the heads of—1st, simple variability; 2nd, polymorphism; 3rd, local forms; 4th, coexisting varieties; 5th, races or subspecies; and 6th, true species.