On the phenomena of variation and geographical distribution as illustrated by the Papilionidæ of the Malayan regionWallace, Alfred Russel
Science
On the phenomena of variation and geographical distribution as illustrated by the Papilionidæ of the Malayan region
Wallace, Alfred Russel
Papilionidae -- Malaysia -- Geographical distribution; Papilionidae -- Malaysia -- Variation
Further east the numbers
decrease, Batchian producing seventeen, and New Guinea only thirteen,
though this number is certainly too small, owing to our present
imperfect knowledge of that great island.
In estimating these numbers I have had the usual difficulty to
encounter, of determining what to consider species and what varieties.
The Malayan region, consisting of a large number of islands of generally
great antiquity, possesses, compared to its actual area, a great number
of distinct forms, often indeed distinguished by very slight characters,
but in most cases so constant in large series of specimens, and so
easily separable from each other, that I know not on what principle we
can refuse to give them the name and rank of species. One of the best
and most orthodox definitions is that of Pritchard, the great
ethnologist, who says, that “_separate origin and distinctness of race,
evinced by a constant transmission of some characteristic peculiarity of
organization_,” constitutes a species. Now leaving out the question of
“origin,” which we cannot determine, and taking only the proof of
separate origin, “_the constant transmission of some characteristic
peculiarity of organization_,” we have a definition which will compel us
to neglect altogether the _amount_ of difference between any two forms,
and to consider only whether the differences that present themselves are
_permanent_. The rule, therefore, I have endeavoured to adopt is, that
when the difference between two forms inhabiting separate areas seems
quite constant, when it can be defined in words, and when it is not
confined to a single peculiarity only, I have considered such forms to
be species. When, however, the individuals of each locality vary among
themselves, so as to cause the distinctions between the two forms to
become inconsiderable and indefinite, or where the differences, though
constant, are confined to one particular only, such as size, tint, or a
single point of difference in marking or in outline, I class one of the
forms as a variety of the other.
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