Phylogeny of the Waxwings and Allied Birds — John Shaqi
Phylogeny of the Waxwings and Allied BirdsArvey, M. Dale (Martin Dale)
Science
Phylogeny of the Waxwings and Allied Birds
Arvey, M. Dale (Martin Dale)
Bombycillidae -- Phylogeny; Waxwings -- Phylogeny
The centrum of _Dulus_ in posterior aspect presents the appearance of
a simple shield; little of the indentation seen in _Bombycilla_ is
present. The spinous portion is plain, with no constriction nor
terminal enlargement in the neck. The centrum in _Phainopepla_ is
similar to that in _Dulus_, but has a small expansion at the base of
the spine, the entire centrum being wider in proportion to its
over-all size than in any of the other species mentioned previously.
The centrum in _Ptilogonys_ shows great width, and the spine is in a
large expanded tip as in _Bombycilla_. The lateral edges of the
centrum in _P. cinereus_ are "winged" and in two separate halves;
whereas the centrum of _P. caudatus_ is fairly plain, its
specialization being reflected primarily in breadth and flatness. In
cross section of the centrum, _Phainoptila_ is similar to
_Phainopepla_, although, in the former, the bone is smaller in
proportion to the size of the animal, and the lateral wings are more
angular than in _Phainopepla_.
[Illustration: Figs. 36-42. Pygostyles in lateral view of five
genera of Bombycillidae. x 2.
36. _Phainoptila m. melanoxantha_, sex?, MNH no. 26493, 15 mi.
SE Cartago, Costa Rica.
37. _Ptilogonys caudatus_, male, MNH no. 24492, 15 mi. SE Cartago,
Costa Rica.
38. _Phainoptila nitens_, male, MNH no. 24754, Pima Co., Arizona.
39. _Ptilogonys cinereus_, female, Louisiana State University
no. 297, Xilitla Region, San Luis Potosi, Mexico.
40. _Dulus dominicus_, female, USNM no. 292652, Don Don, Haiti.
41. _Bombycilla cedrorum_, male, MNH no. 15331, Bexar Co., Texas.
42. _Bombycilla garrula_, sex?, USNM no. 223895, Bozeman, Montana.]
In specialization for muscle attachment, the centra of the pygostyles
of the Ptilogonatinae have more area for muscle attachment than do the
centra in the Bombycillinae and Dulinae; the centrum is wide, the
spinous portion is long, and the bone is flattened anteriorly. The
most generalized pygostyle is in _Phainoptila_, and that of _Dulus_
differs only slightly. In _Bombycilla_ the pygostyle is
proportionately small, but is complex in shape; there is seemingly not
the need for greatly expanded areas since the caudal muscles are less
specialized in this genus.
_Sternum._--The sternum in Bombycillids is typically passerine in
general shape and in having a long and deep carina or sternal crest.
The caudal process of the bone is broad, with the terminal ends
flattened, forming dorsally a graceful V-shaped outline, whereas the
outline of the posterior end of the sternum is broad and convex.
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