Phylogeny of the Waxwings and Allied BirdsArvey, M. Dale (Martin Dale)
Science
Phylogeny of the Waxwings and Allied Birds
Arvey, M. Dale (Martin Dale)
Bombycillidae -- Phylogeny; Waxwings -- Phylogeny
In lateral aspect, the carina is deeper in _Bombycilla_ than in other
genera of the family, and is deepest in _B. garrula_. In this species,
the manubrium is more extended and comparatively larger than in the
other species of the family. The anterior edge of the keel forms the
sharpest angle in _B. cedrorum_. In _Dulus_, the keel is moderately
deep, the manubrium short, and there is a distinct indented curve
between the manubrium and the anterior angle of the keel.
In ventral aspect the lateral processes of the sternum tend to flare
outwards in adult Ptilogonatines on almost the same plane as the rest
of the bone, whereas in _Bombycilla_ and _Dulus_ the same process is
closer to the body of the sternum. In _Bombycilla_ the xiphoid process
is more dorsal in position than in other species in the family, and in
_Dulus_ an upward curve is very noticeable. The process in these two
genera is narrower than in the Ptilogonatinae, and lacks the heavy
distal terminal enlargement which is apparent in _Ptilogonys_.
_Relative Lengths of Bones._--In instances where the animals being
compared are obviously different in over-all size, it is useful to
express the size of a given part in relation to some other part of the
same individual organism if the aim is to obtain clues as to
differences in functions of the parts being compared. Differences in
actual lengths of corresponding bones in two kinds of animals often,
of course, reflect only the difference in over-all size of the
animals. Consequently, the relative size of the part is expressed as a
percentage in this paper. In computing a percentage it is well, of
course, to select some relatively stable part of the animal to use as
a denominator in the mathematical expression that yields the
percentage. The thoracic region of the vertebral column is thought to
be such a part. For example, the length of the humerus divided by the
length of the thoracic region yields, in _Phainopepla_ and
_Ptilogonys_, respective percentages of .84 and .85. These are roughly
the same, whereas the actual lengths of the humeri are 2.21 and 2.39
cm.
Table 4. Lengths of Leg Bones in cm.
=========================+=======+=============+=================
Species | Femur | Tibiotarsus | Tarsometatarsus
-------------------------+-------+-------------+-----------------
Ptilogonys caudatus | 2.04 | 3.10 | 1.94
Ptilogonys cinereus | 1.89 | 2.90 | 1.77
Phainopepla nitens | 1.76 | 2.78 | 1.72
Phainoptila melanoxantha | 2.43 | 3.77 | 2.58
Dulus dominicus | 2.09 | 3.34 | 2.09
Bombycilla garrula | 2.32 | 3.46 | 1.99
Bombycilla cedrorum | 1.92 | 2.95 | 1.64
-------------------------+-------+-------------+-----------------
Table 5. Leg-trunk Ratios (in percent)
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