The “floating cells” in the stomach pockets and in the ampulla, described by Conant, I believe are in part derived from the endothelial cells of the ampulla. That a portion of them may arise from the ovary, as Conant explains, I do not doubt; I have, further, found a mass of floating cells in a small Charybdea quite as Conant describes for Tripedalia (his Fig. 71). In this Charybdea, however, I could find no traces of any ovary. Conant speaks of larger and smaller floating cells, and that the smaller ones are also found in the males. This latter fact agrees with what I have suggested, that some of the floating cells arise in the ampulla. My chief reasons for my supposition, however, are the following: I find globules of the secretion of the ampulla cells in some of the floating cells and also scattered loosely among them (Fig. 19). These globules in and among the floating cells have the same general appearance and a similar staining capacity as the secretion in the ampulla cells. Again, in spaces within some of the ampulla cells I find bodies resembling the floating cells with lumps of the secretion within them (Fig. 18). The conclusion, therefore, lies near that some of the floating cells originate within the cells of the ampulla, engulf within them some of the secretion, and are then expelled into the lumen of the ampulla. Better said, perhaps, they represent portions of the ampulla cells with some of the secretion. I also found several instances in which a floating cell had the appearance of being expelled from an ampulla cell. Conant suggests for a similar observation that the cells were about to be swallowed by the ampulla cells. I believe, however, that my finding a secretion similar to that within the cells of the ampulla, in some of the floating cells, as also bodies very much like them and filled with secretion within the ampulla cells, together with Conant’s finding floating cells in males, and finally the observation that the floating cells are usually quite dilapidated, never showing a healthy cell structure--all this leads me to conclude that some of the floating cells originate from the ampulla cells, and that they have a nutrient function in distributing the secretion. This is quite the reverse of what Conant supposed,--that they were taken in as nourishment by the ampulla cells. I also find what appears to be a secretion in the endoderm of the tentacles of both Charybdea and Tripedalia, and believe this is another source of the floating cells. (See below, under tentacles.)