As I have said above, it is in the cases nearest to hand that the
problem may be most effectively studied. Comparison between forms from
dissimilar situations contributes something; but it is by a close
examination of the behaviour, especially the genetic behaviour, of
familiar species when living in the presence of their nearest allies
that the most direct light on the problem is to be obtained. I cannot
understand the attitude of those who, contemplating such facts as this
examination elicits, can complacently declare that specific difference
is a mere question of degree. With the spread of evolutionary ideas to
speak much of the fixity of species has become unfashionable, and yet
how striking and inscrutable are the manifestations of that fixity!
Consider the group of species composing the _agrestis_ section of the
genus _Veronica_, namely _Tournefortii_, _agrestis_, and _polita_.
These three grow side by side in my garden, as they do in suitable
situations over a vast area of the temperate regions. I have for years
noticed them with some care and become familiar with their distinctions
and resemblances. Never is there any real doubt as to the identity of
any plant. The species show some variability, but I have never seen
one which assumed any of the distinguishing features of the others. A
glance at the fruits decides at once to which species a plant belongs. I
find it impossible to believe that the fixity of these distinctions is
directly dependent on their value as aids in the struggle for existence.
The mode of existence of the three forms in so far as we can tell is
closely similar. By whatever standard we reckon systematic affinity I
suppose we shall agree that these species come very near indeed to each
other. Bentham even takes the view that _polita_ is a mere variety of
_agrestis_.
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