Now in such cases as this it has been argued that the specific
features of the several types have been separately developed in as
many distinct localities, and that their present association is due
to subsequent redistribution. Of these Veronicas indeed we know that
one, _Tournefortii_ (= _Buxbaumii_) is as a matter of fact a recent
introduction from the east.[8] But this course of argument leads to
still further difficulties. For if it is true that the peculiarities
of the several species have been perfected and preserved on account
of their survival-value to their possessors, it follows that there
must be many ways of attaining the same result. But since sufficient
adaptation may be ensured in so many ways, the disappearance of the
common parent of these forms is difficult to understand. Obviously it
must have been a plant very similar in general construction to its
modern representatives. Like them it must have been an annual weed, with
an organisation conformable to that mode of life. Why then, after having
been duly perfected for that existence should it have been entirely
superseded in favour of a number of other distinct contrivances for
doing the same thing, and--if a gradual transition be predicated--not
only by them, but by each intermediate stage between them and the
original progenitor? Surely the obvious inference from such facts is
that the burden cast upon the theory of gradual selection is far greater
than it can bear; that adaptation is not in practice a very close fit,
and that the distinctions between these several species of Veronica have
not arisen on account of their survival-value but rather because none
of their diversities was so damaging as to lead to the extermination
of its possessor. When we see these various Veronicas each rigidly
reproducing its parental type, all comfortably surviving in competition
with each other, are we not forced to the conclusion that _tolerance_
has as much to do with the diversity of species as the stringency of
Selection? Certainly these species owe their continued existence to the
fact that they are each good enough to live, but how shall we refer the
distinctions between them directly or indirectly to the determination of
Natural Selection?
The control of Selection is loose while the conformity to specific
distinction is often very strict and precise, and no less so even when
several closely related species co-exist in the same area and in the
same circumstances.
The theory of Selection fails at exactly the point where it was devised
to help: _Specific_ distinction.
Public-domain text, read in full here on John Shaqi.
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