The asymmetrical, or to speak more precisely, the disorderly, mingling
of the colours in the somatic parts is thus an indication of a similarly
disorderly mixing of the factors for those colours in the germ-tissues,
so that some of the gametes bear enough of the colour-factors to make a
self-coloured plant, while others bear so little that the plant to which
they give rise is a patchwork. If this view is correct we may extend it
so far as to consider whether the fineness or coarseness of the mixture
visible in the flowers or leaves may not give an indication of the
degree to which the factors are subdivided among the germ-cells. We know
very little about the genetic properties of striped varieties. In both
_Antirrhinum_ and _Mirabilis_ it has been found that the striped may
occasionally and irregularly throw self-coloured plants, and therefore
the striping cannot be regarded simply as a recessive character. On the
other hand in _Primula Sinensis_ there are well-known flaked varieties
which ordinarily at least breed true. Whether these ever throw selfs I
do not know, but if they do it must be quite exceptionally. The power
of these flaked plants to breed true is, I suspect, connected with the
fact that in their flowers the coloured and white parts are _intimately_
mixed, this intimate mixture thus being an indication of a similarly
intimate mixture in the germ-cells. It would be important to ascertain
whether self-fertilised seed from the occasional flowers in which the
colour has run together to join a large patch gives more self-coloured
plants than the intimately flaked flowers do.
The next fact may eventually prove of great importance. We have seen
that in bud-sports the differentiation is of the same nature as
that between pure types, and also that in the sporting plant this
differentiation is distributed without any reference to the plant's
axis, or any other consideration of symmetry. Now among the germ-cells
of a Mendelian hybrid exactly such characters are being distributed
allelomorphically, and there again we have strong evidence for believing
that the distribution obeys no pattern. For example, we can in the case
of seeds still _in situ_ perceive how the characters were distributed
among the germ-cells, and there is certainly no obvious pattern
connecting them, nor can we suppose that there is an actual pattern
obscured.
Public-domain text, read in full here on John Shaqi.
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