Of this one illustration is especially curious. Individual plants of
the same species are, as regards the decussations of their leaves and
in other respects, _either rights or lefts_. The fact is not emphasized
in modern botany and is in some danger of being forgotten. When, as
in the flowers of Arum, some _Gladioli_, _Exacum_, _St. Paulia_, or
the fruits of _Loasa_, rights and lefts occur on the same stem,
they come off alternately. But if, as in the seedlings of Barley the
twist of the first leaf be examined, it will be seen to be either a
right-or left-handed screw. An ear of barley, say a two-row barley,
is a definitely symmetrical structure. The seeds stand in their
envelopes back to back in definite positions. Each has its organs
placed in perfectly definite places. _If these seeds were buds_ their
differentiations would be grouped into a common plan. One might expect
that the differentiations of these embryos would still fall into the
pattern; but they do not, and so far as I have tested them, any one
may be a right or a left, just as each may carry any of the Mendelian
allelomorphs possessed by the parent plant, without reference to the
differentiation of any other seed. The fertilisation may be responsible,
but our experience of the allelomorphic characters suggest that the
irregularity is in the egg-cells themselves.[19]
_Germ cells thus differ from somatic cells in the fact that their
differentiations are outside the geometrical order which governs
the differentiation of the somatic cells._ I can think of possible
exceptions, but I have confidence that the rule is true and I regard it
as of great significance.
The old riddle, what is an individual, finds at least a partial solution
in the reply that an individual is a group of parts differentiated in a
geometrically interdependent order. With the germ-cell a new geometrical
order, with independent polarity is almost if not quite always, begun,
and with this geometrical independence the power of rejuvenescence may
possibly be associated.
Public-domain text, read in full here on John Shaqi.
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