Upon this basis the first subdivision of the Radiolaria was made by Johannes Müller, who recognised three groups:--"I. _Thalassicolla_, without receptacle, naked or with spicules; II. _Polycystina_, with a siliceous receptacle; III. _Acanthometra_, without receptacle, but with siliceous radial spines" (L. N. 12, p. 16). 106. _The Ectolithia and Entolithia (Extracapsular and Intracapsular Skeletons)._--The relation of the skeleton to the central capsule in the Radiolaria is very various in many respects; in the first instance two great groups, _Ectolithia_ and _Entolithia_ (see note A), may be distinguished topographically by mere external observation; in the former the skeleton lies entirely outside the central capsule; in the latter, partially at all events, within it. The _Ectolithia_, with a completely extracapsular skeleton, include all NASSELLARIA and PHÆODARIA, as well as a great part of the SPUMELLARIA (all #Collodaria# and the most archaic forms of #Sphærellaria#); the _Entolithia_, on the other hand, in which the skeleton lies partly within, partly without the central capsule, include all ACANTHARIA and the majority of the SPUMELLARIA (most #Sphærellaria#, see note B). A. The difference between Ectolithia and Entolithia was applied in my Monograph in 1862 (p. 222) to separate the Monocyttaria into two main groups. The arrangement was, however, quite artificial, being contrary to the natural relations of the larger groups, as was shown seventeen years later by the discovery of the different structural relations of the central capsule. B. Among the ACANTHARIA, which all possess primitively an intracapsular and centrogenous skeleton, the remarkable _Cenocapsa_ (Pl. 133, fig. 11), seems to furnish the single exception; in it the skeleton consists of a simple spherical shell which encloses the concentric central capsule. The exception is, however, only apparent; the twenty perspinal pores of the shell show that they were originally in connection with twenty centrogenous acanthin spines, and that those have disappeared by retrograde metamorphosis.