Research methods in ecologyClements, Frederic E. (Frederic Edward)
Science
Research methods in ecology
Clements, Frederic E. (Frederic Edward)
Plant ecology; Plant ecology -- Methodology
=245. Arrangement.= The arrangement of species within each formation
herbarium is based upon the structure of the vegetation. The primary
groupings are made with reference to time of appearance and abundance;
when definite zones, associations, or layers are present, they must
likewise be taken into account. In the Colorado collection, the first
division is into three aspects based upon the period of flowering
(_aspectus vernalis_, _aestivalis_, _autumnalis_). Within each aspect,
the species are arranged with respect to abundance in the groups,
facies, principal species, and secondary species. Each group is placed
in an ordinary manila cover, which bears a printed label indicating the
aspect and the group. The species labels give, in addition to the name,
date, and place of collection, the phyad or vegetation form, the
geographical area, the rank of the species, the aspect, and the
formation. To these may well be added data concerning migration
contrivances, seed production, pollination, period of flowering, etc.
The photographs are mounted on the usual herbarium sheets, and placed in
the proper order in the various groups, and a similar disposition is
made of quadrat and transect charts, and such physical factor summaries
as seem desirable.
=246. Succession herbaria.= The arrangement of formation herbaria may
follow the classification of formations with respect to character,
region, or development. The first is the most convenient for purposes of
instruction, and has distinct advantages in permitting a close
comparison of the vegetation of different habitats. The second basis,
which is the one used in the _Herbaria Formationum Coloradensium_, is
peculiarly adapted to mountain vegetation in which the zones are usually
very distinct. The arrangement of herbaria in a developmental series,
however, is the most logical and the most illuminating, since the
structure of the ultimate formations is not only made plain, but the
stages in their development are also laid bare. Such succession herbaria
are the natural outgrowth of formational ones. Indeed, the latter should
be made merely the starting point for these in all regions where the
causes which bring about successions are active. Where weathering is
still an important factor, as in mountains, the initial and intermediate
formations which lead to the final grassland or forest are often in
evidence. After a formation herbarium of each stage has been made in the
way indicated, a succession herbarium is obtained merely by arranging
the various herbaria in the sequence of the developmental stages. Thus,
in the Colorado collection, the subalpine formations are arranged
according to altitude in the following series: (1) the pine formation,
(2) the gravel slide formation, (3) the half gravel slide formation, (4)
the aspen formation, (5) the balsam-spruce formation, (6) the
spruce-pine formation, (7) the meadow thicket formation, (8) the brook
bank formation.
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