Research methods in ecologyClements, Frederic E. (Frederic Edward)
Science
Research methods in ecology
Clements, Frederic E. (Frederic Edward)
Plant ecology; Plant ecology -- Methodology
=296. Succession in alluvial soils.= Alluvial soils are fluvial when
laid down by streams and rivers, and litoral when washed up by the waves
or tides. They are formed when any obstacle retards the movement of the
water, decreasing its carrying power, and causing the deposit of part or
all of its load. They consist of more or less rounded, finely comminuted
particles, mingled with organic matter and detritus. Alluvial deposits
are especially frequent at the mouth of streams and rivers, on their
terraces and flood plains, and along silting banks as compared with the
erosion banks of meanders. The filling of ponds by the erosion due to
surface drainage, and of lakes by the deposition of the loads of streams
that enter them, results in the formation of new alluvium. A similar
phenomenon occurs along coasts, where bays and inlets are slowly
converted into marshes in consequence of being shallowed by the material
washed in by the waves and tides. Such paludal deposits are invariably
salt water or brackish. Contrasted with these, which are uniformly black
in consequence of the large amount of organic matter present, are the
sandbars and beaches, which, though due to the same agents, are light
grey or white in color, because of the constant leaching by the waves.
Two kinds of alluvial deposits may accordingly be distinguished: (1)
those black with organic matter, and little disturbed by water, and (2)
those of a light color, which are constantly swept by the waves. The
successions corresponding to these are radically different. In the
first, the pioneer vegetation is hydrophytic, consisting largely of
amphibious plants. The pioneer stages retard the movement of the water
more and more, and correspondingly hasten the deposition of its load.
The marsh bed slowly rises in consequence, and finally the marsh begins
to dry out, passing first into a wet meadow, and then into a meadow of
the normal type. A notable exception to this sequence occurs when the
swamp contains organic matter or salts in excess, in which case the
vegetation consists indefinitely of swamp xerophytes, or halophytes. The
first vegetation on fresh water sandbars is xerophytic, or, properly,
dissophytic, unless they remain water-swept, and the ultimate stages of
their successions are mesophytic woodlands composed of water-loving
genera, _Populus_, _Salix_, etc. It seems certain, however, that these
will finally give way to longer-lived hardwoods. Maritime sandbars and
beaches are always saline, and their successions run their short course
of development entirely within the group of halophytes, unless the
retreat of the sea or freshwater floods change the character of the
soil. The chemical action of underground waters also produces new soils,
which might be classed as alluvial. These soils are essentially rock
deposits, travertine, silicious sinter, etc., made by iron and lime
springs and by geysers, and they must be changed by decomposition into
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