Research methods in ecologyClements, Frederic E. (Frederic Edward)
Science
Research methods in ecology
Clements, Frederic E. (Frederic Edward)
Plant ecology; Plant ecology -- Methodology
=28. Factors.= The factors of a habitat are water-content, humidity,
light, temperature, soil, wind, precipitation, pressure, altitude,
exposure, slope, surface (cover), and animals. To these should be added
gravity and polarity, which are practically uniform for all habitats,
and may, in consequence, be ignored in this treatise. Length of season,
while it plays an important part in vegetation, is clearly a complex and
is to be treated under its constituents. Of the factors given, all are
regularly found in each habitat, though some are not constantly present.
The first five, water-content, humidity, light, temperature, and soil
are the most important, and any one may well serve as a basis for
grouping habitats into particular classes with reference to quantity. As
will be pointed out later, however, water-content and light furnish the
most striking differences between habitats, and offer the best means of
classification. As habitats are inseparable from the formations which
they bear, the discussion of the kinds of habitats is reserved for
chapter IV.
CLASSIFICATION OF FACTORS
=29. The nature of factors.= The factors of a habitat are arranged in
two groups according to their nature: (1) physical, (2) biotic. In the
strict sense, the physical factors constitute the habitat proper, and
are the real causative forces. No habitat escapes the influence of
biotic factors, however, as the formation always reacts upon it, and the
influence of animals is usually felt in some measure. Physical factors
are further grouped into (1) climatic and (2) edaphic, with respect to
source, or, better, the medium in which they are found. Climatic, or
atmospheric factors are humidity, light, temperature, wind, pressure,
and precipitation. Axiomatically, the stimuli which they produce are
especially related to the leaf. Edaphic or soil factors are confined to
the soil, as the term denotes, and are immediately concerned with the
functions of the root. Water-content is by far the most important of
these; the others are soil composition (nutrient-content), soil
temperature, altitude, slope, exposure, and surface. The last four are
of a more general character than the others, and are usually referred to
as physiographic factors. Cover, when dead, might well be placed among
these also, but as it is little different from the living cover in
effect, it seems most logical to refer it to biotic factors.
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