Research methods in ecologyClements, Frederic E. (Frederic Edward)
Science
Research methods in ecology
Clements, Frederic E. (Frederic Edward)
Plant ecology; Plant ecology -- Methodology
=30. The influence of factors.= While the above classification is both
obvious and convenient, a more logical and intimate grouping may be made
upon the influence which the factor exerts. On this basis, factors are
divided into (1) direct, (2) indirect, and (3) remote. Direct factors
are those which act directly upon an important function of the plant and
produce a formative effect: for example, an increase in humidity
produces an immediate decrease in transpiration. They are water-content,
humidity, and light. Other factors have a direct action: thus
temperature has an immediate influence upon respiration and probably
assimilation also, but it is not structurally formative. Wind has a
direct mechanical effect upon woody plants, but it does not fall within
our definition. Indirect factors are those that affect a formative
function of the plant through another factor; thus a change in
temperature causes a change in humidity and this in turn calls forth a
change in transpiration; or, a change in soil texture increases the
water-content, and this affects the imbibition of the root-hairs.
Indirect factors, then, are temperature, wind, pressure, precipitation,
and soil composition. Remote factors are, for the most part,
physiographic and biotic: they require at least two other factors to act
as middlemen. Altitude affects plants through pressure, which modifies
humidity, and hence transpiration. Slope determines in large degree the
run-off during a rainstorm, thus affecting water-content and the amount
of water absorbed. Earthworms and plant parts change the texture of the
soil, and thereby the water-content. Indirect factors often exert a
remote influence also, as may be seen in the effect which temperature
and wind have in increasing evaporation from the soil, and thus reducing
the water-content. This distinction between factors may seem
insufficiently grounded. In this event, it should be noted that it
centers the effects of all factors upon the three direct ones,
water-content, humidity, and light. If it further be recalled that these
are the only factors which produce qualitative structural changes, and
that the classification of ecads and formations is based upon them, the
validity of the distinction is clear.
THE DETERMINATION OF FACTORS
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