Researches on the Visual Organs of the TrilobitesLindström, Gustaf
Science
Researches on the Visual Organs of the Trilobites
Lindström, Gustaf
Trilobites
[Footnote 6: BEECHER Natural Classification of the Trilobites, p. 191,
pretends that in Olenellus and Holmia real sutures »in a condition of
symphysis» occur. He seems to deny the facial sutures and to accept as
»real sutures» the »internal sutures» described by HOLM in Olenellus
Kjerulfi. It is highly doubtful if these interior elevated lines are
to be regarded as sutures. They are indeed no sutures, but in reality
elevated linear ridges, inclosing, as it were, narrow canals. The real
sutures known, the facial sutures, never form elevated lines, be it
on the outside of the head or on the inside. Probably these lines are
derived from some now unknown interior organization and it may be fit
to remind of the somewhat similar though more numerous linear canals of
the branchiæ on the interior surface of the great head shield of Apus.
(ZADDACH De Apodis cancriformis Anatome ... pl. II fig. 1) or what
HUXLEY (Anatomy Invertebr. Animals p. 281) calls the convoluted »shell
gland» in the carapace. A quite different structure is what I suppose
to be the incipient facial suture in Holmia.]
[Footnote 7: WALCOTT in 10th Ann. Rept. U. S. Geol. Survey, pl. 82 f.
2.]
[Illustration: Larva of Olenellus asaphoides, chiefly according to FORD
and WALCOTT.]
1. Hypothetical figure, based on fig. 2. represents a stage
preceding the next. There are three pair of pleura and three
segments of the rhachis.
2. Copy of WALCOTT'S fig. 5 in »Bullet. 1886», pl. XVII. Three
pleura at right, two at left, the larger formed through fusion
of the two hindmost in fig. 1.
3. Copy of WALCOTT'S fig. 6: the second and third pleura have been
united into one.
4. Hypothetical figure as a further development, following upon
the stage represented in fig. 3. Fusion completed between all
posterior plectra (2, 3, 4), thus forming a single large pleuron
composed of the three mentioned.
5. Copy of FORD'S fig. 1, pl. IV, Amer. Journ. Science 1877,
enlarged to the same size as Walcott's. First sign of the
pygidium.
6. Copy of FORD'S fig. 2 (1877) enlarged. The pygidium has been
added to the head shield.
7. Copy of FORD'S fig. 3, enlarged.
8. Copy of FORD'S fig. 1 page 251 in Am. Journ. Oct. 1881, enlarged.
9. Copy of FORD'S fig. 2 page 251. Amer. Journ. Science 1881.
Slightly enlarged.
10. Copy of FORD'S figure 5, pl. IV, Amer. Journ. Sci. 1877,
slightly enlarged.
Already LINNARSSON observed the second ridge as he tells in a paper
where he describes Olen. Kjerulfi for the first time.[8] It has been
called an »ornamental» spine, but in the following we shall learn what
it really is. This ridge in connection with its spine has not been
observed in any other of the Olenellidæ, at least not in their adult
stage.
[Footnote 8: Öfversigt Vet. Ak. Förhandl. 1871, tafl. XVI fig. 1.]
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